
生物多样性 ›› 2026, Vol. 34 ›› Issue (4): 25438. DOI: 10.17520/biods.2025438 cstr: 32101.14.biods.2025438
刘昊1,2,3(
), 张玉霄4(
), 刘冰1,2,3(
), 李飞飞5,6(
), 马洪峥7(
), 覃海宁1,2,3, 李德铢8(
), 陈文俐1,2,3,*(
)(
)
收稿日期:2025-11-03
接受日期:2026-01-17
出版日期:2026-04-20
发布日期:2026-05-06
通讯作者:
*E-mail: chenwl@ibcas.ac.cn
基金资助:
Hao Liu1,2,3(
), Yuxiao Zhang4(
), Bing Liu1,2,3(
), Feifei Li5,6(
), Hongzheng Ma7(
), Haining Qin1,2,3, Dezhu Li8(
), Wenli Chen1,2,3,*(
)(
)
Received:2025-11-03
Accepted:2026-01-17
Online:2026-04-20
Published:2026-05-06
Contact:
*E-mail: chenwl@ibcas.ac.cn
Supported by:摘要:
通过全面收集、整理中国禾本科植物物种信息, 本文更新了中国禾本科植物物种名录, 系统分析了禾本科植物多样性的分布特点, 总结了禾本科植物标本的保存和采集现状。物种数据统计发现: (1)中国现有禾本科植物10亚科37族283属2,304种, 其中本土物种数包括10亚科34族232属1,978种(含12个特有属和1,028个特有种), 仅次于菊科, 位列中国种子植物第二大科。在本土物种中, 早熟禾亚科(Pooideae)含74属746种, 竹亚科(Bambusoideae)含46属712种, 是物种数最丰富的两个亚科; 簕竹属(Bambusa, 99种)和披碱草属(Elymus, 96种)是物种数最丰富的两个大属。外来物种涉及8亚科20族109属326种, 其中栽培植物占比69%。中国本土禾本科植物的属、种多样性中心位于西南及华南地区, 集中分布了全国92%的本土属和76%的本土种。(2)中国现有国家重点保护禾本科植物21属29种, 省级重点保护禾本科植物15属16种, 受威胁物种31属54种。(3)排名前15位的标本馆其馆藏禾本科标本数合计占全国禾本科植物标本的73%, 这些标本馆主要隶属中国科学院各机构; 现有禾本科植物标本采集多集中于大属和广布种, 而寡种属和狭域分布的特有种采集相对不足。本研究可为禾本科植物的分类、系统学、生物多样性保护和资源可持续利用等方面提供基础数据。
刘昊, 张玉霄, 刘冰, 李飞飞, 马洪峥, 覃海宁, 李德铢, 陈文俐 (2026) 中国禾本科植物多样性及其物种名录. 生物多样性, 34, 25438. DOI: 10.17520/biods.2025438.
Hao Liu, Yuxiao Zhang, Bing Liu, Feifei Li, Hongzheng Ma, Haining Qin, Dezhu Li, Wenli Chen (2026) Diversity and checklist of grasses (Poaceae) in China. Biodiversity Science, 34, 25438. DOI: 10.17520/biods.2025438.
| 主要支系 Major clades | 亚科 Subfamilies | 族 Tribes | 属 Genera | 物种 Species | |||
|---|---|---|---|---|---|---|---|
| N | A | N | A | N | A | ||
| Basal Clade | 原禾亚科 Pharoideae | 1 (1) | 0 | 1 (3) | 0 | 1 (12) | 0 |
| BOP | 稻亚科 Oryzoideae | 1 (4) | 2 | 5 (19) | 3 | 12 (117) | 5 |
| BOP | 竹亚科 Bambusoideae | 2 (3) | 2 | 46 (136) | 23 | 712 (1,698) | 66 |
| BOP | 早熟禾亚科 Pooideae | 12 (15) | 6 | 74 (219) | 37 | 746 (4,126) | 123 |
| PACMAD | 三芒草亚科 Aristidoideae | 1 (1) | 0 | 2 (3) | 0 | 12 (367) | 0 |
| PACMAD | 黍亚科 Panicoideae | 7 (14) | 4 | 68 (242) | 30 | 351 (3,325) | 93 |
| PACMAD | 芦竹亚科 Arundinoideae | 3 (3) | 1 | 4 (14) | 2 | 7 (36) | 2 |
| PACMAD | 澳禾亚科 Micrairoideae | 2 (3) | 1 | 4 (9) | 1 | 22 (192) | 1 |
| PACMAD | 扁芒草亚科 Danthonioideae | 1 (1) | 1 | 2 (19) | 2 | 4 (292) | 2 |
| PACMAD | 虎尾草亚科 Chloridoideae | 4 (5) | 3 | 26 (121) | 11 | 111 (1,603) | 34 |
| 合计 Total | 34 (54) | 20 | 232 (789) | 109 | 1,978 (11,783) | 326 | |
| 去重统计 Unique count | 10 | 37 | 283 | 2,304 | |||
表1 中国禾本科植物的多样性组成
Table 1 The diversity composition of Poaceae in China
| 主要支系 Major clades | 亚科 Subfamilies | 族 Tribes | 属 Genera | 物种 Species | |||
|---|---|---|---|---|---|---|---|
| N | A | N | A | N | A | ||
| Basal Clade | 原禾亚科 Pharoideae | 1 (1) | 0 | 1 (3) | 0 | 1 (12) | 0 |
| BOP | 稻亚科 Oryzoideae | 1 (4) | 2 | 5 (19) | 3 | 12 (117) | 5 |
| BOP | 竹亚科 Bambusoideae | 2 (3) | 2 | 46 (136) | 23 | 712 (1,698) | 66 |
| BOP | 早熟禾亚科 Pooideae | 12 (15) | 6 | 74 (219) | 37 | 746 (4,126) | 123 |
| PACMAD | 三芒草亚科 Aristidoideae | 1 (1) | 0 | 2 (3) | 0 | 12 (367) | 0 |
| PACMAD | 黍亚科 Panicoideae | 7 (14) | 4 | 68 (242) | 30 | 351 (3,325) | 93 |
| PACMAD | 芦竹亚科 Arundinoideae | 3 (3) | 1 | 4 (14) | 2 | 7 (36) | 2 |
| PACMAD | 澳禾亚科 Micrairoideae | 2 (3) | 1 | 4 (9) | 1 | 22 (192) | 1 |
| PACMAD | 扁芒草亚科 Danthonioideae | 1 (1) | 1 | 2 (19) | 2 | 4 (292) | 2 |
| PACMAD | 虎尾草亚科 Chloridoideae | 4 (5) | 3 | 26 (121) | 11 | 111 (1,603) | 34 |
| 合计 Total | 34 (54) | 20 | 232 (789) | 109 | 1,978 (11,783) | 326 | |
| 去重统计 Unique count | 10 | 37 | 283 | 2,304 | |||
| 族 Tribes | 物种数 No. of species | % | 属 Genera | 物种数 No. of species | % |
|---|---|---|---|---|---|
| 1. 北美箭竹族 Arundinarieae | 511 | 25.8 | 1. 簕竹属 Bambusa | 99 | 5.0 |
| 2. 早熟禾族 Poeae | 367 | 18.6 | 2. 披碱草属 Elymus | 96 | 4.9 |
| 3. 高粱族 Andropogoneae | 201 | 10.2 | 3. 箭竹属 Fargesia | 94 | 4.8 |
| 4. 簕竹族 Bambuseae | 201 | 10.2 | 4. 玉山竹属 Yushania | 91 | 4.6 |
| 5. 小麦族 Triticeae | 185 | 9.4 | 5. 早熟禾属 Poa | 82 | 4.1 |
| 6. 黍族 Paniceae | 109 | 5.5 | 6. 刚竹属 Phyllostachys | 63 | 3.2 |
| 7. 针茅族 Stipeae | 88 | 4.4 | 7. 碱茅属 Puccinellia | 55 | 2.8 |
| 8. 狗牙根族 Cynodonteae | 57 | 2.9 | 8. 羊茅属 Festuca | 55 | 2.8 |
| 9. 雀麦族 Bromeae | 51 | 2.6 | 9. 雀麦属 Bromus | 51 | 2.6 |
| 10. 画眉草族 Eragrostideae | 36 | 1.8 | 10. 方竹属 Chimonobambusa | 48 | 2.4 |
| 合计 Total | 1,806 | 91.4 | 合计 Total | 734 | 37.2 |
表2 中国本土禾本科植物物种数排名前十位的族和属
Table 2 Top 10 tribes and genera by number of native Poaceae species in China
| 族 Tribes | 物种数 No. of species | % | 属 Genera | 物种数 No. of species | % |
|---|---|---|---|---|---|
| 1. 北美箭竹族 Arundinarieae | 511 | 25.8 | 1. 簕竹属 Bambusa | 99 | 5.0 |
| 2. 早熟禾族 Poeae | 367 | 18.6 | 2. 披碱草属 Elymus | 96 | 4.9 |
| 3. 高粱族 Andropogoneae | 201 | 10.2 | 3. 箭竹属 Fargesia | 94 | 4.8 |
| 4. 簕竹族 Bambuseae | 201 | 10.2 | 4. 玉山竹属 Yushania | 91 | 4.6 |
| 5. 小麦族 Triticeae | 185 | 9.4 | 5. 早熟禾属 Poa | 82 | 4.1 |
| 6. 黍族 Paniceae | 109 | 5.5 | 6. 刚竹属 Phyllostachys | 63 | 3.2 |
| 7. 针茅族 Stipeae | 88 | 4.4 | 7. 碱茅属 Puccinellia | 55 | 2.8 |
| 8. 狗牙根族 Cynodonteae | 57 | 2.9 | 8. 羊茅属 Festuca | 55 | 2.8 |
| 9. 雀麦族 Bromeae | 51 | 2.6 | 9. 雀麦属 Bromus | 51 | 2.6 |
| 10. 画眉草族 Eragrostideae | 36 | 1.8 | 10. 方竹属 Chimonobambusa | 48 | 2.4 |
| 合计 Total | 1,806 | 91.4 | 合计 Total | 734 | 37.2 |
| 本文 This study | FOC Vol. 22 | 参考文献 References | |||
|---|---|---|---|---|---|
| 新属 New genera | |||||
| 1. 异燕麦属 Helictochloa* | 隶属于山燕麦属 Genus previously included in Helictotrichon | Romero-Zarco, | |||
| 2. 痕花黍属 Hildaea | 隶属于距花黍属 Genus previously included in Ichnanthus | Silva et al., | |||
| 3. 纪如竹属 Hsuehochloa* | 隶属于悬竹属 Genus previously included in Ampelocalamus | Zhang et al., | |||
| 4. 以礼竹属 Kengiochloa* | 隶属于矢竹属 Genus previously included in Pseudosasa | Tong et al., | |||
| 5. 新芨芨草属(新拟) Neotrinia* | 隶属于芨芨草属 Genus previously included in Achnatherum | Nobis et al., | |||
| 6. 雷文竹属 Ravenochloa* | 隶属于箬竹属 Genus previously included in Indocalamus | Zhang et al., | |||
| 7. 三毛草属 Sibirotrisetum* | 隶属于穗三毛草属 Genus previously included in Trisetum | Barberá et al., | |||
| 8. 华赤竹属 Sinosasa* | 隶属于赤竹属 Genus previously included in Sasa | Qin et al., | |||
| 9. 头梨竹属 Stapletonia | 新属 New genus | Dash et al., | |||
| 10. 坚丛草属(新拟) Tenaxia | 隶属于扁芒草属 Genus previously included in Danthonia | Linder et al., | |||
| 11. 棉花竹属 Tongpeia* | 隶属于箭竹属 Genus previously included in Fargesia | Stapleton, | |||
| 12. 斑茅属(新拟) Tripidium | 隶属于甘蔗属 Genus previously included in Saccharum | Valdés & Scholz, | |||
| 13. 绿燕麦属(新拟) Trisetopsis | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 14. 茨燕麦属(新拟) Tzveleviochloa | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 15. 高异燕麦属 ×Trisetopsotrichon | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 恢复属 Resurrected genera | |||||
| 1. 类早熟禾属 Arctopoa* | 隶属于早熟禾属 Genus previously included in Poa | Soreng et al., | |||
| 2. 曲芒发草属 Avenella | 隶属于发草属 Genus previously included in Deschampsia | Soreng et al., | |||
| 3. 巴山木竹属 Bashania* | 隶属于北美箭竹属 Genus previously included in Arundinaria | Vorontsova et al., | |||
| 4. 短穗竹属 Brachystachyum | 隶属于业平竹属 Genus previously included in Semiarundinaria | Zhang et al., | |||
| 5. 小沿沟草属 Catabrosella | 隶属于拟沿沟草属 Genus previously included in Colpodium | Soreng et al., | |||
| 6. 肿节竹属 Clavinodum | 隶属于少穗竹属 Genus previously included in Oligostachyum | Niu et al., | |||
| 7. 假拟沿沟草 Paracolpodium | 隶属于拟沿沟草属 Genus previously included in Colpodium | Soreng et al., | |||
| 8. 茅针属 Patis* | 隶属于芨芨草属 Genus previously included in Achnatherum | Romaschenko et al., | |||
| 9. 冷箭竹属 Sarocalamus | 隶属于北美箭竹属 Genus previously included in Arundinaria | Vorontsova et al., | |||
| 10. 华箬竹属 Sasamorpha | 隶属于箬竹属 Genus previously included in Indocalamus | Vorontsova et al., | |||
| 归并属 Merged genera | |||||
| 1. 凤头黍属 Acroceras (=Setiacis*) | 刺毛头黍属 Setiacis | Soreng et al., | |||
| 2. 蒺藜草属 Cenchrus* (=Pennisetum*) | 狼尾草属 Pennisetum | Chemisquy et al., | |||
| 3. 鼠尾粟属 Sporobolus* (=Crypsis) | 隐花草属 Crypsis | Peterson et al., | |||
| 4. 尾稃草属 Urochloa* (=Brachiaria*) | 臂形草属 Brachiaria | Soreng et al., | |||
| 新记录属 Newly recorded genera | |||||
| 1. 穗燕麦属(新拟) Acrospelion | 新组合 New combination | Barberá et al., | |||
| 2. 多枝竹属 Holttumochloa* | 新物种 New species | Zhou et al., | |||
| 3. 五须芒属 Pentapogon | 新组合 New combination | Peterson et al., | |||
| 4. 雅竹属 Temochloa* | 新物种 New species | Cai et al. | |||
表3 本文与Flora of China (FOC) Vol. 22 (Chen et al., 2006)以及《中国生物物物种名录(2025版)》植物类群数据库(Liu et al., 2025)本土属级变化的比较
Table 3 Generic-level taxonomic changes in different native genera of Poaceae in China compared with Flora of China (FOC) Vol. 22 (Chen et al., 2006) and China Checklist of Higher Plants in the Catalogue of Life China 2025 Annual Checklist (Liu et al., 2025)
| 本文 This study | FOC Vol. 22 | 参考文献 References | |||
|---|---|---|---|---|---|
| 新属 New genera | |||||
| 1. 异燕麦属 Helictochloa* | 隶属于山燕麦属 Genus previously included in Helictotrichon | Romero-Zarco, | |||
| 2. 痕花黍属 Hildaea | 隶属于距花黍属 Genus previously included in Ichnanthus | Silva et al., | |||
| 3. 纪如竹属 Hsuehochloa* | 隶属于悬竹属 Genus previously included in Ampelocalamus | Zhang et al., | |||
| 4. 以礼竹属 Kengiochloa* | 隶属于矢竹属 Genus previously included in Pseudosasa | Tong et al., | |||
| 5. 新芨芨草属(新拟) Neotrinia* | 隶属于芨芨草属 Genus previously included in Achnatherum | Nobis et al., | |||
| 6. 雷文竹属 Ravenochloa* | 隶属于箬竹属 Genus previously included in Indocalamus | Zhang et al., | |||
| 7. 三毛草属 Sibirotrisetum* | 隶属于穗三毛草属 Genus previously included in Trisetum | Barberá et al., | |||
| 8. 华赤竹属 Sinosasa* | 隶属于赤竹属 Genus previously included in Sasa | Qin et al., | |||
| 9. 头梨竹属 Stapletonia | 新属 New genus | Dash et al., | |||
| 10. 坚丛草属(新拟) Tenaxia | 隶属于扁芒草属 Genus previously included in Danthonia | Linder et al., | |||
| 11. 棉花竹属 Tongpeia* | 隶属于箭竹属 Genus previously included in Fargesia | Stapleton, | |||
| 12. 斑茅属(新拟) Tripidium | 隶属于甘蔗属 Genus previously included in Saccharum | Valdés & Scholz, | |||
| 13. 绿燕麦属(新拟) Trisetopsis | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 14. 茨燕麦属(新拟) Tzveleviochloa | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 15. 高异燕麦属 ×Trisetopsotrichon | 隶属于山燕麦属 Genus previously included in Helictotrichon | Wölk & Röser, | |||
| 恢复属 Resurrected genera | |||||
| 1. 类早熟禾属 Arctopoa* | 隶属于早熟禾属 Genus previously included in Poa | Soreng et al., | |||
| 2. 曲芒发草属 Avenella | 隶属于发草属 Genus previously included in Deschampsia | Soreng et al., | |||
| 3. 巴山木竹属 Bashania* | 隶属于北美箭竹属 Genus previously included in Arundinaria | Vorontsova et al., | |||
| 4. 短穗竹属 Brachystachyum | 隶属于业平竹属 Genus previously included in Semiarundinaria | Zhang et al., | |||
| 5. 小沿沟草属 Catabrosella | 隶属于拟沿沟草属 Genus previously included in Colpodium | Soreng et al., | |||
| 6. 肿节竹属 Clavinodum | 隶属于少穗竹属 Genus previously included in Oligostachyum | Niu et al., | |||
| 7. 假拟沿沟草 Paracolpodium | 隶属于拟沿沟草属 Genus previously included in Colpodium | Soreng et al., | |||
| 8. 茅针属 Patis* | 隶属于芨芨草属 Genus previously included in Achnatherum | Romaschenko et al., | |||
| 9. 冷箭竹属 Sarocalamus | 隶属于北美箭竹属 Genus previously included in Arundinaria | Vorontsova et al., | |||
| 10. 华箬竹属 Sasamorpha | 隶属于箬竹属 Genus previously included in Indocalamus | Vorontsova et al., | |||
| 归并属 Merged genera | |||||
| 1. 凤头黍属 Acroceras (=Setiacis*) | 刺毛头黍属 Setiacis | Soreng et al., | |||
| 2. 蒺藜草属 Cenchrus* (=Pennisetum*) | 狼尾草属 Pennisetum | Chemisquy et al., | |||
| 3. 鼠尾粟属 Sporobolus* (=Crypsis) | 隐花草属 Crypsis | Peterson et al., | |||
| 4. 尾稃草属 Urochloa* (=Brachiaria*) | 臂形草属 Brachiaria | Soreng et al., | |||
| 新记录属 Newly recorded genera | |||||
| 1. 穗燕麦属(新拟) Acrospelion | 新组合 New combination | Barberá et al., | |||
| 2. 多枝竹属 Holttumochloa* | 新物种 New species | Zhou et al., | |||
| 3. 五须芒属 Pentapogon | 新组合 New combination | Peterson et al., | |||
| 4. 雅竹属 Temochloa* | 新物种 New species | Cai et al. | |||
图2 中国禾本科植物馆藏和采集标本数据统计。(A)标本数前15位的标本馆馆藏量(标本馆信息见附录4); (B)属级标本采集数; (C)物种级标本采集数。
Fig. 2 Statistics of herbarium holdings and specimen collection for Poaceae in China. (A) Holdings of the top 15 herbaria by specimen count (herbarium information is listed in Appendix 4); (B) Collection numbers at the generic level; (C) Collection numbers at the species level.
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