生物多样性 ›› 2019, Vol. 27 ›› Issue (11): 1228-1235.DOI: 10.17520/biods.2019111
赵国华1,2,王莹2,商辉2,周喜乐3,王爱华4,李玉峰5,王晖1,刘保东6,严岳鸿2,*()
收稿日期:
2019-03-29
接受日期:
2019-10-08
出版日期:
2019-11-20
发布日期:
2020-01-17
通讯作者:
严岳鸿
基金资助:
Guohua Zhao1,2,Ying Wang2,Hui Shang2,Xile Zhou3,Aihua Wang4,Yufeng Li5,Hui Wang1,Baodong Liu6,Yuehong Yan2,*()
Received:
2019-03-29
Accepted:
2019-10-08
Online:
2019-11-20
Published:
2020-01-17
Contact:
Yan Yuehong
摘要:
蕨类植物孢子表面的纹饰形态复杂多样, 在分类学和古生物学研究中具有重要意义, 目前探索祖先性状演化常用形态学与系统发育学结合的方法。本文选取一回羽叶铁线蕨类植物为材料, 用扫描电子显微镜观察其孢子形态, 用5个叶绿体基因序列构建系统树, 用最大似然法和最大简约法来分析孢子性状的演化。结果表明: 一回羽叶铁线蕨类的孢子纹饰有5种类型, 即光滑、粗糙、颗粒、疣状及瘤状; 这一类群的祖先孢子纹饰有很大的可能是瘤状纹饰; 疣状纹饰可能是由瘤状纹饰演化而来; 孢子纹饰由简单向复杂演化, 越进化的物种其孢子纹饰越复杂。
赵国华, 王莹, 商辉, 周喜乐, 王爱华, 李玉峰, 王晖, 刘保东, 严岳鸿. 祖先性状重建法揭示铁线蕨属植物孢子表面纹饰的形态多样性及其演化[J]. 生物多样性, 2019, 27(11): 1228-1235.
Guohua Zhao, Ying Wang, Hui Shang, Xile Zhou, Aihua Wang, Yufeng Li, Hui Wang, Baodong Liu, Yuehong Yan. Ancestral state reconstruction reveals the diversity and evolution of spore ornamentation in Adiantum (Pteridaceae)[J]. Biodiv Sci, 2019, 27(11): 1228-1235.
物种 Species | 标本号及采集地 Vouchers and locality | 赤道轴 × 极轴 Equatorial diam × polar diam (µm) | 孢子纹饰 Spore ornamentation | Position in |
---|---|---|---|---|
A. latifolium | Zxl9882. 泰国曼谷植物标本馆 Bangkok Herbarium, Bangkok, Thailand | 33.5 × 27.0 | 粗糙 Scabrate | 1A |
扇叶铁线蕨 A. flabellulatum | Zxl10018. 中国广东南岭国家森林公园 Nanling National Forest Park, Guangdong, China | 45.5 × 31.2 | 颗粒 Granulate | 1B |
团羽铁线蕨 A. capillus-junonis | Zxl9959. 中国广东南岭国家森林公园 Nanling National Forest Park, Guangdong, China | 46.5 × 43.0 | 光滑 Psilate | 1C |
半月形铁线蕨 A. philippense | Yan12405, Yan12086. 中国云南元江; Zxl9839. 泰国彭世洛府屯萨朗峦国家公园 Yan12405, Yan12086. Yuanjiang, Yunnan, China; Zxl9839. Thung Salaeng Luang National Park, Phitsanulok, Thailand | 33.4 × 25.7 | 粗糙 Scabrate | 1D |
孟连铁线蕨 A. menglianense | Yan12415, Yan12409. 中国云南元江 Yuanjiang, Yunnan, China | 31.5 × 26.6 | 颗粒 Granulate | 1E |
翅柄铁线蕨 A. soboliferum | Yan11393. 中国海南昌江 Changjiang, Hainan, China | 34.0 × 30.5 | 颗粒 Granulate | 1F |
A. zollingeri | Zxl9812, Zxl9911, ZXL9851. 泰国碧武里府康卡沾国家公园 Kaeng Krachan National Park, Phetchaburi, Thailand | 34.0 × 27.3 | 瘤状 Tuberculate | 1G |
鞭叶铁线蕨 A. caudatum | Yan12408. 中国云南元江 Yuanjiang, Yunnan, China | 40.6 × 33.5 | 瘤状 Tuberculate | 1H |
白垩铁线蕨 A. gravesii | WYG393. 中国贵州贞丰 Zhenfeng, Guizhou, China | 51.5 × 36.1 | 瘤状 Tuberculate | 1I |
小铁线蕨 A. mariesii | Zxl9685. 中国湖南桑植 Sangzhi, Hunan, China | 36.8 × 30.1 | 疣状 Verrucate | 1J |
仙霞铁线蕨 A. juxtapositum | Yan1205071. 中国广东仁化 Renhua, Guangdong, China | 60.4 × 45.7 | 疣状 Verrucate | 1K |
普通铁线蕨 A. edgewothii | Yan12412. 中国云南元江 Yuanjiang, Yunnan, China | 30.0 × 25.6 | 瘤状 Tuberculate | 1L |
假鞭叶铁线蕨 A. malesianum | Yan12416, Yan12404. 中国云南元江 Yuanjiang, Yunnan, China | 38.8 × 32.8 | 瘤状 Tuberculate | 1M |
梅山铁线蕨 A. × meishanianum | Yan12411. 中国云南元江 Yuanjiang, Yunnan, China | - | 瘤状 Tuberculate | 1N |
苍山铁线蕨 A. sinicum | Yan112601, Yan12414. 中国云南元江 Yuanjiang, Yunnan, China | 31.7 × 29.1 | 瘤状 Tuberculate | 1O |
哀牢山铁线蕨 A. × ailaoshanense | Yan12413, Yan12410. 中国云南元江 Yuanjiang, Yunnan, China | - | 瘤状 Tuberculate | 1P |
表1 铁线蕨属植物采集信息及孢子形态
Table 1 The locality information and spore morphology in Adiantum
物种 Species | 标本号及采集地 Vouchers and locality | 赤道轴 × 极轴 Equatorial diam × polar diam (µm) | 孢子纹饰 Spore ornamentation | Position in |
---|---|---|---|---|
A. latifolium | Zxl9882. 泰国曼谷植物标本馆 Bangkok Herbarium, Bangkok, Thailand | 33.5 × 27.0 | 粗糙 Scabrate | 1A |
扇叶铁线蕨 A. flabellulatum | Zxl10018. 中国广东南岭国家森林公园 Nanling National Forest Park, Guangdong, China | 45.5 × 31.2 | 颗粒 Granulate | 1B |
团羽铁线蕨 A. capillus-junonis | Zxl9959. 中国广东南岭国家森林公园 Nanling National Forest Park, Guangdong, China | 46.5 × 43.0 | 光滑 Psilate | 1C |
半月形铁线蕨 A. philippense | Yan12405, Yan12086. 中国云南元江; Zxl9839. 泰国彭世洛府屯萨朗峦国家公园 Yan12405, Yan12086. Yuanjiang, Yunnan, China; Zxl9839. Thung Salaeng Luang National Park, Phitsanulok, Thailand | 33.4 × 25.7 | 粗糙 Scabrate | 1D |
孟连铁线蕨 A. menglianense | Yan12415, Yan12409. 中国云南元江 Yuanjiang, Yunnan, China | 31.5 × 26.6 | 颗粒 Granulate | 1E |
翅柄铁线蕨 A. soboliferum | Yan11393. 中国海南昌江 Changjiang, Hainan, China | 34.0 × 30.5 | 颗粒 Granulate | 1F |
A. zollingeri | Zxl9812, Zxl9911, ZXL9851. 泰国碧武里府康卡沾国家公园 Kaeng Krachan National Park, Phetchaburi, Thailand | 34.0 × 27.3 | 瘤状 Tuberculate | 1G |
鞭叶铁线蕨 A. caudatum | Yan12408. 中国云南元江 Yuanjiang, Yunnan, China | 40.6 × 33.5 | 瘤状 Tuberculate | 1H |
白垩铁线蕨 A. gravesii | WYG393. 中国贵州贞丰 Zhenfeng, Guizhou, China | 51.5 × 36.1 | 瘤状 Tuberculate | 1I |
小铁线蕨 A. mariesii | Zxl9685. 中国湖南桑植 Sangzhi, Hunan, China | 36.8 × 30.1 | 疣状 Verrucate | 1J |
仙霞铁线蕨 A. juxtapositum | Yan1205071. 中国广东仁化 Renhua, Guangdong, China | 60.4 × 45.7 | 疣状 Verrucate | 1K |
普通铁线蕨 A. edgewothii | Yan12412. 中国云南元江 Yuanjiang, Yunnan, China | 30.0 × 25.6 | 瘤状 Tuberculate | 1L |
假鞭叶铁线蕨 A. malesianum | Yan12416, Yan12404. 中国云南元江 Yuanjiang, Yunnan, China | 38.8 × 32.8 | 瘤状 Tuberculate | 1M |
梅山铁线蕨 A. × meishanianum | Yan12411. 中国云南元江 Yuanjiang, Yunnan, China | - | 瘤状 Tuberculate | 1N |
苍山铁线蕨 A. sinicum | Yan112601, Yan12414. 中国云南元江 Yuanjiang, Yunnan, China | 31.7 × 29.1 | 瘤状 Tuberculate | 1O |
哀牢山铁线蕨 A. × ailaoshanense | Yan12413, Yan12410. 中国云南元江 Yuanjiang, Yunnan, China | - | 瘤状 Tuberculate | 1P |
图1 一回羽叶铁线蕨类孢子形态。 (A) A. latifolium; (B)扇叶铁线蕨; (C)团羽铁线蕨; (D)半月形铁线蕨; (E)孟连铁线蕨; (F)翅柄铁线蕨; (G) A. zollingeri; (H)鞭叶铁线蕨; (I)白垩铁线蕨; (J)小铁线蕨; (K)仙霞铁线蕨; (L)普通铁线蕨; (M)假鞭叶铁线蕨; (N)梅山铁线蕨; (O)苍山铁线蕨; (P)哀牢山铁线蕨。
Fig. 1 Spore morphology in once-pinnate maidenhair ferns (Adiantum). (A) A. latifolium; (B) A. flabellulatum; (C) A. capillus-junonis; (D) A. philippense; (E) A. menglianense; (F) A. soboliferum; (G) A. zollingeri; (H) A. caudatum; (I) A. gravesii; (J) A. mariesii; (K) A. juxtapositum; (L) A. edgewothii; (M) A. malesianum; (N) A. × meishanianum; (O) A. sinicum; (P) A. × ailaoshanense.
图2 铁线蕨属孢子大小。黄色代表白垩铁线蕨系, 蓝色代表鞭叶铁线蕨系, 黑色代表外类群。
Fig. 2 Spore size of Adiantum. Yellow shapes indicate A. Ser. Gravesiana. Blue shapes indicate A. Ser. Caudata. Black shapes indicate outgroup.
图3 5个叶绿体序列rbcL、atpB、atpA、trnL-F、rps4-trnS联合构建系统发生树, 数字代表支持率(ML/MP/BI)。*: ML和MP = 100%, BI = 1.0。
Fig. 3 The phylogenetic tree constructed based on five chloroplast markers (rbcL, atpB, atpA, trnL-F, rps4-trnS). Numbers on branches are support values (ML/MP/BI). *, ML, MP = 100%, and BI = 1.0.
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