生物多样性 ›› 2021, Vol. 29 ›› Issue (9): 1256-1264. DOI: 10.17520/biods.2021119
收稿日期:
2021-03-31
接受日期:
2021-07-10
出版日期:
2021-09-20
发布日期:
2021-08-19
通讯作者:
张鹗
作者简介:
* E-mail: zhange@ihb.ac.cn基金资助:
Received:
2021-03-31
Accepted:
2021-07-10
Online:
2021-09-20
Published:
2021-08-19
Contact:
E Zhang
摘要:
赣江是长江的主要支流之一, 淡水鱼类多样性极其丰富, 但是目前对于该河流的鱼类多样性还未有充分的了解。本研究于2016年9月至2017年8月对赣江鱼类进行了实地调查, 结合历史文献资料并参考最新的分类学成果, 更新了赣江淡水鱼类名录。结果表明, 赣江共有淡水鱼类180种, 隶属于12目31科93属。其中, 土著鱼类174种, 外来鱼类6种。赣江鱼类以鲤形目为主; 鲤科种类最多, 其次是鲿科。更新的物种名录中, 包含23个新记录土著种, 其中有5个未被描述的新种; 有36个历史资料记载的种类被剔除出物种名录; 25个物种发生了分类地位的变更。此外, 本次调查有28种鱼类未采集到, 说明这些鱼类的种群数量正在急剧下降, 而这些鱼类分别属于洄游性、流水性和产漂流性卵等独特生态类型, 说明赣江鱼类多样性受到了严重的人为干扰。本研究所给出的赣江鱼类的更新物种名录, 可为今后的赣江鱼类多样性保护提供理论基础。
王子彤, 张鹗 (2021) 赣江鱼类物种更新名录. 生物多样性, 29, 1256-1264. DOI: 10.17520/biods.2021119.
Zitong Wang, E Zhang (2021) An updated species checklist of freshwater fishes from the Ganjiang River. Biodiversity Science, 29, 1256-1264. DOI: 10.17520/biods.2021119.
[1] |
Akai Y, Arai R (1998) Rhodeus sinensis, a senior synonym of R. lighti and R. uyekii (Acheilognathinae, Cyprinidae). Ichthyological Research, 45, 105-110.
DOI URL |
[2] |
An CT, Zhang E, Shen JZ (2020) Sarcocheilichthys vittatus, a new species of gudgeon (Teleostei: Cyprinidae) from the Poyang Lake basin in Jiangxi Province, South China. Zootaxa, 4768, 201-220.
DOI URL |
[3] | Banarescu PM (1997) The status of some nominal genera of Eurasian Cyprinidae (Osteichthyes, Cypriniformes). Revue Roumaine de Biologie, Série de Biologie Animale, 42, 19-30. |
[4] | Berg LS (2004) Freshwater Fishes of the USSR and Adjacent Countries. Publishing House of Academy of Science of USSR, Moskva. |
[5] | Bogutskaya NG, Naseka AM (2004) Catalogue of Agnathans and Fishes of Fresh and Brackish Waters of Russia with Comments on Nomenclature and Taxonomy. KMK Scientific Press Ltd, Moscow. |
[6] |
Bohlen J, Šlechtová V (2017) Leptobotia micra, a new species of loach (Teleostei: Botiidae) from Guilin, southern China. Zootaxa, 4250, 90-100.
DOI PMID |
[7] |
Cao L, Zhang E, Zang CX, Cao WX (2016) Evaluating the status of China's continental fish and analyzing their causes of endangerment through the red list assessment. Biodiversity Science, 24, 598-610. (in Chinese with English abstract)
DOI URL |
[曹亮, 张鹗, 臧春鑫, 曹文宣 (2016) 通过红色名录评估研究中国内陆鱼类受威胁现状及其成因. 生物多样性, 24, 598-610.]
DOI |
|
[8] | Cao WX (2017) Aquatic environment protection induced by the cascade hydropower development in the upper Yangtze River. Technology and Economy of Changjiang, (1), 25-30. (in Chinese) |
[曹文宣 (2017) 长江上游水电梯级开发的水域生态保护问题. 长江技术经济, 1, 25-30. | |
[9] | Cao YH, Liao FC, Wu YA (2012) Aquatic Fauna of Xiangjiang River. Hunan Science and Technology Publishing House, Changsha. (in Chinese) |
[曹英华, 廖伏初, 伍远安 (2012) 湘江水生动物志. 湖南科学技术出版社, 长沙.] | |
[10] | Chen YY (1998) Fauna Sinica·Osteichthyes·Cypriniformes.II. Science Press, Beijing. (in Chinese) |
[陈宜瑜 (1998) 中国动物志·硬骨鱼纲·鲤形目(中). 科学出版社, 北京.] | |
[11] | Chu XL (1989) The Fishes of Yunnan. Part 1. Science Press, Beijing. (in Chinese) |
[褚新洛 (1989) 云南鱼类志(上册). 科学出版社, 北京.] | |
[12] | Chu XL (1999) Fauna Sinica·Osteichthyes·Siluriformes. Science Press, Beijing. (in Chinese) |
[褚新洛 (1999) 中国动物志·硬骨鱼纲·鲇形目. 科学出版社, 北京.] | |
[13] | Ding RH (1994) The Fishes of Sichuan, China. Sichuan Publishing House of Science and Technology, Chengdu. (in Chinese) |
[丁瑞华 (1994) 四川鱼类志. 四川科学技术出版社, 成都.] | |
[14] |
Dudgeon D, Arthington AH, Gessner MO, Kawabata ZI, Knowler DJ, Lévêque C, Naiman RJ, Prieur-Richard AH, Soto D, Stiassny MLJ, Sullivan CA (2006) Freshwater biodiversity: Importance, threats, status and conservation challenges. Biological Reviews, 81, 163-182.
PMID |
[15] | Gao TY, Xie D, Peng ND, Li C, Zhang SP, Luo JZ, Wang CH, Zhao J (2017) Fish community structure and its relationships with environmental factors in the Beijiang River. Freshwater Fisheries, 47(6), 47-55. (in Chinese with English abstract) |
[高天扬, 谢迪, 彭宁东, 李潮, 张少平, 罗锦桢, 王春晖, 赵俊 (2017) 北江鱼类群落结构及其与环境因子的关系. 淡水渔业, 47(6), 47-55.] | |
[16] | Guo Q, Liu XJ, Ao XF, Qin JJ, Wu XP, Ouyang S (2018) Fish diversity in the middle and lower reaches of the Ganjiang River of China: Threats and conservation. PLoS ONE, 13, e205116. |
[17] | Guo ZZ, Liu RL (1995) The fishes of Jiangxi Province. Journal of Nanchang University (Natural Science), 19, 222-232. (in Chinese with English abstract) |
[郭治之, 刘瑞兰 (1995) 江西鱼类的研究. 南昌大学学报(理科版), 19, 222-232.] | |
[18] | Guo ZZ, Zou DL, Liu RL, Pan YH, Ye GY, Liu SP (1964) The investigation report on fish in Poyang Lake. Journal of Nanchang University (Natural Science), (2), 121-130. (in Chinese) |
[郭治之, 邹多录, 刘瑞兰, 潘永浩, 叶桂瑶, 刘世平 (1964) 鄱阳湖鱼类调查报告. 南昌大学学报(理科版), (2), 121-130.] | |
[19] | Hu ML, Wu ZQ, Li Q, Huang LL (2011) Preliminary research on fish species diversity of Ganjiangyuan Nature Reserve in Jiangxi Province. Sichuan Journal of Zoology, 30, 467-470. (in Chinese with English abstract) |
[胡茂林, 吴志强, 李晴, 黄亮亮 (2011) 江西赣江源自然保护区鱼类物种多样性初步研究. 四川动物, 30, 467-470.] | |
[20] |
Huang LL, Wu ZQ, Li JH (2013) Fish fauna, biogeography and conservation of freshwater fish in Poyang Lake basin, China. Environmental Biology of Fishes, 96, 1229-1243.
DOI URL |
[21] | Huang SP, Zhao YH, Chen IS, Shao KT (2017) A new species of Microphysogobio (Cypriniformes: Cyprinidae) from Guangxi Province, Southern China. Zoological Studies, 56, 1-12. |
[22] | Ito T, Hosoya K (2016) Re-examination of the type series of Parazacco spilurus (Teleostei: Cyprinidae). Fishtaxa, 1, 89-93. |
[23] |
Jiang ZG, Gao EH, Zhang E (2012) Microphysogobio nudiventris, a new species of gudgeon (Teleostei: Cyprinidae) from the middle Chang-Jiang (Yangtze River) basin, Hubei Province, South China. Zootaxa, 3586, 211-221.
DOI URL |
[24] | Kottelat M (2006) Fishes of Mongolia: A Check-list of the Fishes Known to Occur in Mongolia with Comments on Systematics and Nomenclature. Environment and Social Development, East Asia and Pacific Region, World Bank, Washington, DC. |
[25] | Kottelat M (2012) Conspectus cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26(Suppl.), 1-199. |
[26] | Kottelat M (2013) The fishes of the inland waters of Southeast Asia: A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology, 27(Suppl.), 1-663. |
[27] | Kottelat M, Freyhof J (2007) Handbook of European Freshwater Fish. Kottelat, Cornol & Freyhof, Berlin. |
[28] |
Ku XY, Peng ZG, Diogo R, He SP (2007) MtDNA phylogeny provides evidence of generic polyphyleticism for East Asian bagrid catfishes. Hydrobiologia, 579, 147-159.
DOI URL |
[29] | Li Q, Wu ZQ, Huang LL, Chen HM, Lu HJ (2008) Fish resources of Qiyunshan Nature Reserve, Jiangxi, China. Acta Zootaxonomica Sinica, 33, 324-329. (in Chinese with English abstract) |
[李晴, 吴志强, 黄亮亮, 陈辉敏, 卢和军 (2008) 江西齐云山自然保护区鱼类资源. 动物分类学报, 33, 324-329.] | |
[30] |
Li S, Chen JK, Wang XM (2016) Global distribution, entry routes, mechanisms and consequences of invasive freshwater fish. Biodiversity Science, 24, 672-685. (in Chinese with English abstract)
DOI URL |
[郦珊, 陈家宽, 王小明 (2016) 淡水鱼类入侵种的分布、入侵途径、机制与后果. 生物多样性, 24, 672-685.]
DOI |
|
[31] | Liu LG, Yang CY, Yang PH, Wang WB, Zou WS, Han Q (2013) Status and diversity of fish resources of Yuanshui River in Hunan Province, China. Oceanologia et Limnologia Sinica, 44, 148-158. (in Chinese with English abstract) |
[刘良国, 杨春英, 杨品红, 王文彬, 邹万生, 韩庆 (2013) 湖南境内沅水鱼类资源现状与多样性分析. 海洋与湖沼, 44, 148-158.] | |
[32] |
Liu XJ, Ao XF, Ning ZT, Hu XY, Wu XP, Ouyang S (2017) Diversity of fish species in Suichuan River and Shushui River and conservation value, China. Environmental Biology of Fishes, 100, 493-507.
DOI URL |
[33] | Liu Y, Lin XT, Sun J, Zhang PF, Chen GZ (2011) Fish community changes in Huizhou segment of Dongjiang River. Chinese Journal of Zoology, 46(2), 1-11. (in Chinese with English abstract) |
[刘毅, 林小涛, 孙军, 张鹏飞, 陈国柱 (2011) 东江下游惠州河段鱼类群落组成变化特征. 动物学杂志, 46(2), 1-11.] | |
[34] | Miao C, Zhu YH, Yuan RB, Zou ZA, Hu ML, Hu SP, Yin H, Yan X (2016) Fish species diversity in Jinpenshan Nature Reserve Zone, Jiangxi Province, in summer. Chinese Journal of Fisheries, 29(4), 28-32. (in Chinese with English abstract) |
[苗春, 祝于红, 袁荣斌, 邹志安, 胡茂林, 胡绍平, 尹宏, 严星 (2016) 江西金盆山自然保护区夏季鱼类物种多样性初步调查. 水产学杂志, 29(4), 28-32.] | |
[35] | Ng HH, Freyhof J (2007) Pseudobagrus nubilosus, a new species of catfish from central Vietnam (Teleostei: Bagridae), with notes on the validity of Pelteobagrus and Pseudobagrus. Ichthyological Exploration of Freshwaters, 18, 9-16. |
[36] | Ng HH, Kottelat M (2007) The identity of Tachysurus sinensis Lacepede, 1803, with the designation of a neotype (Teleostei: Bagridae) and notes on the identity of T. fulvidraco (Richardson, 1845). Electronic Journal of Ichthyology, 3, 35-54. |
[37] | Regan CT (1908) XXIV. Descriptions of new freshwater fishes from China and Japan. Annals & Magazine of Natural History, 1, 149-153. |
[38] |
Sakai H, Watanabe K, Goto A (2020) A revised generic taxonomy for Far East Asian minnow Rhynchocypris and dace Pseudaspius. Ichthyological Research, 67, 330-334.
DOI URL |
[39] |
Schönhuth S, Vukić J, Šanda R, Yang L, Mayden RL (2018) Phylogenetic relationships and classification of the Holarctic family Leuciscidae (Cypriniformes: Cyprinoidei). Molecular Phylogenetics and Evolution, 127, 781-799.
DOI PMID |
[40] |
Song XJ, Tang WQ, Zhang Y (2017) Freshwater fish fauna and zoogeographical divisions in the Wuyi-Xianxialing Mountains of eastern China. Biodiversity Science, 25, 1331-1338. (in Chinese with English abstract)
DOI URL |
[宋小晶, 唐文乔, 张亚 (2017) 华东武夷山-仙霞岭地区淡水鱼类区系特征及其动物地理区划. 生物多样性, 25, 1331-1338.]
DOI |
|
[41] |
Song XL, Cao L, Zhang E (2018) Onychostoma brevibarba, a new cyprinine fish (Pisces: Teleostei) from the middle Chang Jiang basin in Hunan Province, South China. Zootaxa, 4410, 147-163.
DOI URL |
[42] | Su N, Li L, Xu ZQ, Xiao D (2012) Status of fish resources in Xiajiang to Nanchang section of Ganjiang River. Journal of Huazhong Agricultural University, 31, 756-764. (in Chinese with English abstract) |
[苏念, 李莉, 徐哲奇, 肖舵 (2012) 赣江峡江至南昌段鱼类资源现状. 华中农业大学学报, 31, 756-764.] | |
[43] | Sun ZW, Ren SJ, Zhang E (2013) Liobagrus chenghaiensis, a new species of catfish (Siluriformes: Amblycipitidae) from Yunnan, South China. Ichthyological Exploration of Freshwaters, 23, 375-384. |
[44] |
Tang QY, Liu HZ, Yang XP, Nakajima T (2005) Molecular and morphological data suggest that Spinibarbus caldwelli (Nichols) (Teleostei: Cyprinidae) is a valid species. Ichthyological Research, 52, 77-82.
DOI URL |
[45] | Tang WQ, Liu HZ, Ma JA, Xiao RC (1993) Effects of Jiangxi Wan'an Dam on Tenualosa reevesii breeding in Ganjiang and the countermeasures. Reservoir Fisheries, (4), 18-19. (in Chinese) |
[唐文乔, 刘焕章, 马经安, 肖仁春 (1993) 江西万安水利枢纽对赣江鲥繁殖的影响及其对策. 水利渔业, (4), 18-19.] | |
[46] | Tian JL (1989) Fish resources in Ganjiang River before the construction of Wan'an Dam and its fishery utilization. Freshwater Fisheries, (1), 33-39. (in Chinese) |
[田见龙 (1989) 万安大坝截流前赣江鱼类调查及渔业利用意见. 淡水渔业, (1), 33-39.] | |
[47] | Tu FY, Li JX, Han WJ, Huang XF (2016) Species diversity and protection of fishes in Jiangxi. Acta Agriculturae Universitatis Jiangxiensis, 38, 975-985. (in Chinese with English abstract) |
[涂飞云, 李久煊, 韩卫杰, 黄晓凤 (2016) 江西省鱼类物种多样性及其保护. 江西农业大学学报, 38, 975-985.] | |
[48] | Wang CY, Huang DC, Hu ML, Liu J, Miao C, Wang T, Jian SQ (2019) Comparative analysis of fish community diversity in two primary tributaries of the lower Ganjiang River. Journal of Hydroecology, 40, 99-106. (in Chinese with English abstract) |
[王朝阳, 黄德才, 胡茂林, 柳俊, 苗春, 王焘, 简少卿 (2019) 赣江下游一级支流鱼类多样性比较分析. 水生态学杂志, 40, 99-106.] | |
[49] |
Wang X, Liu F, Yu D, Liu HZ (2019) Mitochondrial divergence suggests unexpected high species diversity in the opsariichthine fishes (Teleostei: Cyprinidae) and the revalidation of Opsariichthys macrolepis. Ecology and Evolution, 9, 2664-2677.
DOI |
[50] |
Wei FW, Nie YG, Miao HX, Lu H, Hu YB (2014) Advancements of the researches on biodiversity loss mechanisms. Chinese Science Bulletin, 59, 430-437. (in Chinese with English abstract)
DOI URL |
[魏辅文, 聂永刚, 苗海霞, 路浩, 胡义波 (2014) 生物多样性丧失机制研究进展. 科学通报, 59, 430-437.] | |
[51] | Xie ZG, Zhang E, He SP (2001) Study on species validation for Glyptothorax sinense (Regan) and G. fukiensis (Rendahl) with the method of morphometrics. Journal of Huazhong Agricultural University, 20, 169-172. (in Chinese with English abstract) |
[谢仲桂, 张鹗, 何舜平 (2001) 应用形态度量学方法对中华纹胸鮡和福建纹胸鮡物种有效性的研究. 华中农业大学学报, 20, 169-172.] | |
[52] | Yi BL, Zhu ZR (1959) Studies on Culter and Erythroculter fishes of China. Collected Papers of Hydrobiologica, (2), 170-196. (in Chinese) |
[易伯鲁, 朱志荣 (1959) 中国的鲌属和红鲌属鱼类的研究. 水生生物学集刊, (2), 170-196.] | |
[53] |
Yi WJ, Zhang E, Shen JZ (2014) Vanmanenia maculata, a new species of hillstream loach from the Chang-Jiang Basin, South China (Teleostei: Gastromyzontidae). Zootaxa, 3802, 85-97.
DOI URL |
[54] |
Yuan LY, Zhang E (2010) Type locality and identity of Acrossocheilus kreyenbergii (Regan, 1908), a senior synonym of Acrossocheilus cinctus (Lin, 1931)(Teleostei: Cyprinidae). Zootaxa, 2684, 36-44.
DOI URL |
[55] | Yue PQ (2000) Fauna Sinica·Osteichthyes·Cypriniformes. III. Science Press, Beijing. (in Chinese) |
[乐佩琦 (2000) 中国动物志·硬骨鱼纲·鲤形目(下). 科学出版社, 北京.] | |
[56] | Zhang CG (2016) Species Diversity and Distribution of Inland Fishes in China. Science Press, Beijing. (in Chinese) |
[张春光 (2016) 中国内陆鱼类物种与分布. 科学出版社, 北京.] | |
[57] | Zhang E, Xie ZG, Xie CX (2004) Morphological variation between Sinibrama macrops and S. wui, with notes on their validities. Acta Hydrobiologica Sinica, 28, 511-518. (in Chinese with English abstract) |
[张鹗, 谢仲桂, 谢从新 (2004) 大眼华鳊和伍氏华鳊的形态差异及其物种有效性. 水生生物学报, 28, 511-518.] | |
[58] | Zhang JM, Wu ZQ, Hu ML, Zhu RC, Liu BB, Chen YL, Hua Q (2009) Status of fish resources in Xiajiang reach of Middle Ganjiang River. Jiangxi Science, 27, 916-919. (in Chinese with English abstract) |
[张建铭, 吴志强, 胡茂林, 朱日财, 刘彬彬, 陈彦良, 花麒 (2009) 赣江中游峡江段鱼类资源现状. 江西科学, 27, 916-919.] | |
[59] | Zhu L, Yu D, Liu HZ (2020) Zacco sinensis (Cypriniformes: Cyprinidae), a new fish species from Northern China. Sichuan Journal of Zoology, 39, 168-176. (in Chinese with English abstract) |
[朱兰, 俞丹, 刘焕章 (2020) 中国北方鱲属鱼类一新种--中华鱲(鲤形目: 鲤科). 四川动物, 39, 168-176.] | |
[60] | Zhu SQ (1995) Synopsis of Freshwater Fishes of China. Jiangsu Science and Technology Publishing House, Nanjing. (in Chinese) |
[朱松泉 (1995) 中国淡水鱼类检索. 江苏科学技术出版社, 南京.] | |
[61] | Zou DL (1982) Fish fauna of Jiulianshan area, Jiangxi, China. Journal of Jiangxi University (Natural Science), (2), 50-53. (in Chinese) |
[邹多录 (1982) 江西省九连山地区的鱼类及其区系. 江西大学学报(自然科学版), (2), 50-53.] | |
[62] | Zou SZ (2011) Effects of Large Hydraulic Project on Fish and Ecological Environment in the Middle Reach of Gan River. PhD dissertation, Nanchang University, Nanchang. (in Chinese with English abstract) |
[邹淑珍 (2011) 赣江中游大型水利工程对鱼类及其生态环境的影响研究. 博士学位论文, 南昌大学, 南昌.] |
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[10] | 曹焕喜, 罗阿蓉, 牛泽清, 张彦周, 肖晖, 朱朝东. 2021年膜翅目新增分类单元[J]. 生物多样性, 2022, 30(8): 22162-. |
[11] | 王科, 蔡磊. 世界菌物新命名发表概况(2021年)[J]. 生物多样性, 2022, 30(8): 22277-. |
[12] | 林晨, 张冰, 王亮, 杨定. 2021年中国双翅目新分类单元[J]. 生物多样性, 2022, 30(8): 22192-. |
[13] | 王婷, 舒江平, 顾钰峰, 李艳清, 杨拓, 徐洲锋, 向建英, 张宪春, 严岳鸿. 中国石松类和蕨类植物多样性研究进展[J]. 生物多样性, 2022, 30(7): 22381-. |
[14] | 朱瑞良, 马晓英, 曹畅, 曹子寅. 中国苔藓植物多样性研究进展[J]. 生物多样性, 2022, 30(7): 22378-. |
[15] | 杜诚, 刘军, 刘夙, 马金双. 中国植物分类学者的历史与现状[J]. 生物多样性, 2022, 30(7): 22355-. |
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