
生物多样性 ›› 2026, Vol. 34 ›› Issue (5): 25041. DOI: 10.17520/biods.2025041 cstr: 32101.14.biods.2025041
• 研究报告: 生物文化多样性 • 下一篇
收稿日期:2025-01-26
接受日期:2025-10-15
出版日期:2026-05-20
发布日期:2026-07-01
通讯作者:
李仁
基金资助:
Yulu Pu1,2(
), Shishun Zhou2(
), Ren Li2,3,*(
)(
)
Received:2025-01-26
Accepted:2025-10-15
Online:2026-05-20
Published:2026-07-01
Contact:
Ren Li
Supported by:摘要:
香料植物是大航海时代以来的主要贸易商品之一, 具有重要的经济和药用价值。开展香料植物的民族植物学研究, 符合现代社会需求和大健康产业的发展趋势。民族植物学对植物资源的开发和民族地区经济的发展有重要的促进作用, 为有关人类安全和具有药理活性的药物筛选工作提供了前所未有的线索。中国、缅甸、老挝三国热带地区是全球生物多样性研究热点地区之一, 具有丰富的香料植物资源和文化多样性。本研究以中国、缅甸、老挝的傣族、哈尼族、瑶族、老族、克木人、缅族、钦族和克钦族等8个民族利用的香料植物为研究对象, 基于民族植物学原理和多学科方法, 通过1985‒2024年中英文文献检索、野外科考、半结构式访谈、市场和村寨调查, 完成了三国热带香料植物的民族植物学研究。结果显示, 该地区民族利用的香料植物414种(含变种), 分属于63个科和206个属。其用途涵盖药用(300种, 含药食两用74种)、蔬菜(123种)、调味(95种)等, 其利用部位以叶(46.62%)、根(20.77%)为主。利用价值超过0.2的植物大多为药食两用的香料植物。资源来源中野生的有178种(43.00%)、栽培的有185种(44.69%), 这其中仅有12%实现了从野生采集到人工栽培再到市场销售的全过程。中缅、中老、缅老边境地区民族利用的香料植物Jaccard相似性指数为0.20‒0.25, 整体偏低, 表明不同地区民族在长期适应各自生态环境和文化背景下形成了各具特色的植物利用传统, 反映了该地区丰富的文化多样性和民族植物学知识的独特性。此外, 调查发现处于保护或受威胁的物种有52种(12.56%), 经评估和筛选, 其中有15种需要重点研究、保护和利用。本研究完成了全球第一份系统全面展示中国、缅甸、老挝三国热带地区民族利用香料植物的民族植物学编目, 既可为三国香料植物资源在食品、药品和化妆品产业的可持续开发和利用提供知识原型, 也为实施跨国境文化多样性和生物多样性保护, 建设绿色“一带一路”提供基础数据和科学案例。
浦瑀璐, 周仕顺, 李仁 (2026) 中国、缅甸、老挝热带香料植物的民族植物学. 生物多样性, 34, 25041. DOI: 10.17520/biods.2025041.
Yulu Pu, Shishun Zhou, Ren Li (2026) Ethnobotany of tropical spice plants utilized by ethnic groups in China, Myanmar and Laos. Biodiversity Science, 34, 25041. DOI: 10.17520/biods.2025041.
| 国家 Country | 地点 Location | 经纬度 Coordinates (N, E) |
|---|---|---|
| 中国 China | 云南省西双版纳傣族自治州 Xishuangbanna Dai Autonomous Prefecture, Yunnan Province 云南省普洱市 Pu’er City, Yunnan Province | 21°10′-22°40′, 99°55′-101°50′ 22°02′-24°50′, 99°09′-102°19′ |
| 缅甸 Myanmar | 钦邦 Chin State 克钦邦 Kachin State | 20°40′-24°06′, 92°37′-94°09′ 23°27′-28°25′, 96°0′-98°44′ |
| 老挝 Laos | 琅南塔省 Luang Namtha Province 丰沙里省 Phongsali Province 乌多姆赛省 Oudomxay Province | 19°90′-22°50′, 100°10′-103°35′ |
表1 中国、缅甸、老挝热带香料植物重点研究区域
Table 1 Key study areas of tropical spice plants in China, Myanmar and Laos
| 国家 Country | 地点 Location | 经纬度 Coordinates (N, E) |
|---|---|---|
| 中国 China | 云南省西双版纳傣族自治州 Xishuangbanna Dai Autonomous Prefecture, Yunnan Province 云南省普洱市 Pu’er City, Yunnan Province | 21°10′-22°40′, 99°55′-101°50′ 22°02′-24°50′, 99°09′-102°19′ |
| 缅甸 Myanmar | 钦邦 Chin State 克钦邦 Kachin State | 20°40′-24°06′, 92°37′-94°09′ 23°27′-28°25′, 96°0′-98°44′ |
| 老挝 Laos | 琅南塔省 Luang Namtha Province 丰沙里省 Phongsali Province 乌多姆赛省 Oudomxay Province | 19°90′-22°50′, 100°10′-103°35′ |
图2 缅甸钦邦(A1-A4)和克钦邦(B1-B4)民族利用香料植物植物学调查
Fig. 2 Ethnobotanical survey of spice plants used by local people in Chin State (A1-A4) and Kachin State (B1-B4), Myanmar
| 科名 Family name | 物种数 No. of species | 比例 Proportion (%) | 主要生长习性 Main growth habits | 核心用途 Core uses |
|---|---|---|---|---|
| 姜科 Zingiberaceae | 83 | 20.05 | 草本 Herb | 调味、药用 Seasoning, medicinal |
| 菊科 Asteraceae | 51 | 12.32 | 草本、半灌木 Herb, subshrub | 药用、驱蚊 Medicinal, mosquito repellent |
| 唇形科 Lamiaceae | 46 | 11.11 | 草本、灌木 Herb, shrub | 调味、茶饮 Seasoning, tea |
| 芸香科 Rutaceae | 41 | 9.90 | 乔木、灌木 Tree, shrub | 调味、药用 Seasoning, medicinal |
| 樟科 Lauraceae | 26 | 6.52 | 乔木、灌木 Tree, shrub | 调味、驱虫 Seasoning, insect repellent |
| 胡椒科 Piperaceae | 16 | 3.86 | 藤本、半灌木 Vine, subshrub | 调味、药用 Seasoning, medicinal |
| 伞形科 Apiaceae | 16 | 3.86 | 草本 Herb | 药用、蔬菜 Medicinal, vegetable |
表2 中国、缅甸、老挝三国热带香料植物优势科(前7科)
Table 2 The dominant families of tropical spice plants in China, Myanmar and Laos (Top 7)
| 科名 Family name | 物种数 No. of species | 比例 Proportion (%) | 主要生长习性 Main growth habits | 核心用途 Core uses |
|---|---|---|---|---|
| 姜科 Zingiberaceae | 83 | 20.05 | 草本 Herb | 调味、药用 Seasoning, medicinal |
| 菊科 Asteraceae | 51 | 12.32 | 草本、半灌木 Herb, subshrub | 药用、驱蚊 Medicinal, mosquito repellent |
| 唇形科 Lamiaceae | 46 | 11.11 | 草本、灌木 Herb, shrub | 调味、茶饮 Seasoning, tea |
| 芸香科 Rutaceae | 41 | 9.90 | 乔木、灌木 Tree, shrub | 调味、药用 Seasoning, medicinal |
| 樟科 Lauraceae | 26 | 6.52 | 乔木、灌木 Tree, shrub | 调味、驱虫 Seasoning, insect repellent |
| 胡椒科 Piperaceae | 16 | 3.86 | 藤本、半灌木 Vine, subshrub | 调味、药用 Seasoning, medicinal |
| 伞形科 Apiaceae | 16 | 3.86 | 草本 Herb | 药用、蔬菜 Medicinal, vegetable |
| 生长习性 Growth habit | 物种数 No. of species | 比例 Proportion (%) | 生长习性 Growth habit | 物种数 No. of species | 比例 Proportion (%) |
|---|---|---|---|---|---|
| 草本 Herb | 227 | 54.83 | 藤本 Vine | 13 | 3.14 |
| 乔木 Tree | 94 | 22.71 | 木质藤本 Liana | 7 | 1.69 |
| 灌木 Shrub | 55 | 13.29 | 小乔木 Small tree | 5 | 1.21 |
| 半灌木 Subshrub | 13 | 3.14 |
表3 中国、缅甸、老挝三国热带地区民族利用香料植物不同生长习性
Table 3 Different growth habits of tropical spice plants in China, Myanmar and Laos
| 生长习性 Growth habit | 物种数 No. of species | 比例 Proportion (%) | 生长习性 Growth habit | 物种数 No. of species | 比例 Proportion (%) |
|---|---|---|---|---|---|
| 草本 Herb | 227 | 54.83 | 藤本 Vine | 13 | 3.14 |
| 乔木 Tree | 94 | 22.71 | 木质藤本 Liana | 7 | 1.69 |
| 灌木 Shrub | 55 | 13.29 | 小乔木 Small tree | 5 | 1.21 |
| 半灌木 Subshrub | 13 | 3.14 |
| 利用部位 Used part | 物种数 No. of species | 比例 Proportion (%) | 代表物种 Representative species |
|---|---|---|---|
| 叶 Leaf | 193 | 46.62 | 薄荷、蒟叶 Mentha haplocalyx, Piper betle |
| 根 Root | 86 | 20.77 | 姜黄、当归 Curcuma longa, Angelica sinensis |
| 花 Flower | 68 | 16.43 | 菊花、金银花 Chrysanthemum morifolium, Lonicera japonica |
| 果实 Fruit | 60 | 14.49 | 胡椒、花椒 Piper nigrum, Zanthoxylum bungeanum |
| 全草 Whole herb | 42 | 10.14 | 积雪草、鱼腥草 Centella asiatica, Houttuynia cordata |
表4 中国、缅甸、老挝三国热带香料植物主要利用部位统计
Table 4 Statistics on the main used parts of tropical spice plants in China, Myanmar and Laos
| 利用部位 Used part | 物种数 No. of species | 比例 Proportion (%) | 代表物种 Representative species |
|---|---|---|---|
| 叶 Leaf | 193 | 46.62 | 薄荷、蒟叶 Mentha haplocalyx, Piper betle |
| 根 Root | 86 | 20.77 | 姜黄、当归 Curcuma longa, Angelica sinensis |
| 花 Flower | 68 | 16.43 | 菊花、金银花 Chrysanthemum morifolium, Lonicera japonica |
| 果实 Fruit | 60 | 14.49 | 胡椒、花椒 Piper nigrum, Zanthoxylum bungeanum |
| 全草 Whole herb | 42 | 10.14 | 积雪草、鱼腥草 Centella asiatica, Houttuynia cordata |
| 类别 Category | 物种数 No. of species | 比例 Proportion (%) | 主要用途 Main uses |
|---|---|---|---|
| 药用 Medicinal | 300 | 72.46 | 治疗肠胃、风湿、感冒等(含药食两用74种) Treating gastrointestinal disorders, rheumatism, colds, etc. (including 74 medicinal-edible dual-purpose species) |
| 蔬菜 Vegetable | 123 | 29.71 | 日常饮食补充 Daily food supplement (including 74 medicinal-edible dual-purpose species) |
| 调味 Seasoning | 95 | 22.95 | 提升肉类、汤类风味 Enhancing the flavor of meat and soups |
| 驱蚊 Mosquito repellent | 25 | 6.04 | 燃烧、铺放驱蚊虫 Repelling mosquitoes by burning or placing |
| 茶饮 Tea | 16 | 3.86 | 泡水作清凉饮料 Making cool drinks by soaking in water |
表5 中国、缅甸、老挝三国热带香料植物主要用途
Table 5 The main usage of tropical spice plants in China, Myanmar and Laos
| 类别 Category | 物种数 No. of species | 比例 Proportion (%) | 主要用途 Main uses |
|---|---|---|---|
| 药用 Medicinal | 300 | 72.46 | 治疗肠胃、风湿、感冒等(含药食两用74种) Treating gastrointestinal disorders, rheumatism, colds, etc. (including 74 medicinal-edible dual-purpose species) |
| 蔬菜 Vegetable | 123 | 29.71 | 日常饮食补充 Daily food supplement (including 74 medicinal-edible dual-purpose species) |
| 调味 Seasoning | 95 | 22.95 | 提升肉类、汤类风味 Enhancing the flavor of meat and soups |
| 驱蚊 Mosquito repellent | 25 | 6.04 | 燃烧、铺放驱蚊虫 Repelling mosquitoes by burning or placing |
| 茶饮 Tea | 16 | 3.86 | 泡水作清凉饮料 Making cool drinks by soaking in water |
| 类别 Category | 具体方法 Specific methods | 物种数 No. of species | 比例 Proportion (%) | 适用用途 Applicable uses |
|---|---|---|---|---|
| 加热处理 Heat treatment | 煮水喝、做汤、炒吃 Boiling for drinking, making soups, stir-frying | 323 | 78.02 | 药用、蔬菜、调味 Medicinal, vegetable, seasoning |
| 外用处理 External application | 捣碎包敷、涂抹、燃烧 Mashing for compressing, smearing, burning | 105 | 25.36 | 药用、驱蚊 Medicinal, mosquito repellent |
| 生食/凉拌 Raw/eaten cold | 凉拌、生吃、沾剁生 Cold mixing, eating raw, dipping in minced raw meat | 85 | 20.53 | 蔬菜、调味 Vegetable, seasoning |
| 加工利用 Indivece uses | 泡茶、泡酒、染色 Making tea, soaking in wine, dyeing | 73 | 17.63 | 茶饮、染料、药用 Tea, dye, medicinal |
表6 中国、缅甸、老挝三国热带香料植物不同使用方法
Table 6 Different use methods of tropical spice plants in China, Myanmar and Laos
| 类别 Category | 具体方法 Specific methods | 物种数 No. of species | 比例 Proportion (%) | 适用用途 Applicable uses |
|---|---|---|---|---|
| 加热处理 Heat treatment | 煮水喝、做汤、炒吃 Boiling for drinking, making soups, stir-frying | 323 | 78.02 | 药用、蔬菜、调味 Medicinal, vegetable, seasoning |
| 外用处理 External application | 捣碎包敷、涂抹、燃烧 Mashing for compressing, smearing, burning | 105 | 25.36 | 药用、驱蚊 Medicinal, mosquito repellent |
| 生食/凉拌 Raw/eaten cold | 凉拌、生吃、沾剁生 Cold mixing, eating raw, dipping in minced raw meat | 85 | 20.53 | 蔬菜、调味 Vegetable, seasoning |
| 加工利用 Indivece uses | 泡茶、泡酒、染色 Making tea, soaking in wine, dyeing | 73 | 17.63 | 茶饮、染料、药用 Tea, dye, medicinal |
| 来源 Source | 物种数 No of species | 比例 Proportion (%) |
|---|---|---|
| 纯野生 Wild | 178 | 43.00 |
| 栽培和市场有售 Cultivated and commercially available | 65 | 15.70 |
| 野生和市场有售 Wild and commercially available | 51 | 12.32 |
| 野生、栽培和市场有售 Wild, cultivated and commercially available | 50 | 12.08 |
| 纯栽培 Cultivated | 44 | 10.63 |
| 野生和栽培 Wild and cultivated | 26 | 6.28 |
表7 中国、缅甸、老挝民族利用香料植物资源来源组成
Table 7 Source composition of utilized spice plants in China, Myanmar and Laos
| 来源 Source | 物种数 No of species | 比例 Proportion (%) |
|---|---|---|
| 纯野生 Wild | 178 | 43.00 |
| 栽培和市场有售 Cultivated and commercially available | 65 | 15.70 |
| 野生和市场有售 Wild and commercially available | 51 | 12.32 |
| 野生、栽培和市场有售 Wild, cultivated and commercially available | 50 | 12.08 |
| 纯栽培 Cultivated | 44 | 10.63 |
| 野生和栽培 Wild and cultivated | 26 | 6.28 |
| 序号 Serial number | 中文名 Chinese name | 科 Family | 学名 Scientific name | NPLC | TSLCHP | IUCN | UV |
|---|---|---|---|---|---|---|---|
| 1 | 姜状三七 | Araliaceae | Panax zingiberensis | 1 | EN | EN | 0.02 |
| 2 | 香子含笑 | Magnoliaceae | Michelia gioii | 2 | EN | DD | 0.06 |
| 3 | 土沉香 | Thymelaeaceae | Aquilaria sinensis | 2 | VU | VU | 0.02 |
| 4 | 茴香砂仁 | Zingiberaceae | Etlingera yunnanensis | 2 | VU | LC | 0.02 |
| 5 | 紫檀 | Leguminosae | Pterocarpus indicus | - | CR | EN | 0.07 |
| 6 | 毛叶樟 | Lauraceae | Camphora mollifolia | - | EN | EN | 0.03 |
| 7 | 云南石梓 | Lamiaceae | Gmelina arborea | - | VU | LC | 0.20 |
| 8 | 木橘 | Rutaceae | Aegle marmelos | - | VU | NT | 0.14 |
| 9 | 卡西松 | Pinaceae | Pinus kesiya | - | VU | LC | 0.06 |
| 10 | 辣椒砂仁 | Zingiberaceae | Amomum capsiciforme | - | - | CR | 0.08 |
| 11 | 厚叶沉香 | Thymelaeaceae | Aquilaria crassna | - | - | CR | 0.03 |
| 12 | 黄樟 | Lauraceae | Camphora parthenoxylon | - | - | EN | 0.06 |
| 13 | 喙花姜 | Zingiberaceae | Rhynchanthus beesianus | - | EN | - | 0.02 |
| 14 | 毛大叶臭花椒 | Rutaceae | Zanthoxylum myriacanthum var. pubescens | - | VU | - | 0.12 |
| 15 | 毛姜花 | Zingiberaceae | Hedychium villosum | - | VU | - | 0.03 |
表8 筛选和推荐的15种值得进一步深入研究和加强保护的香料植物
Table 8 Fifteen selected spice plants for further research and conservation
| 序号 Serial number | 中文名 Chinese name | 科 Family | 学名 Scientific name | NPLC | TSLCHP | IUCN | UV |
|---|---|---|---|---|---|---|---|
| 1 | 姜状三七 | Araliaceae | Panax zingiberensis | 1 | EN | EN | 0.02 |
| 2 | 香子含笑 | Magnoliaceae | Michelia gioii | 2 | EN | DD | 0.06 |
| 3 | 土沉香 | Thymelaeaceae | Aquilaria sinensis | 2 | VU | VU | 0.02 |
| 4 | 茴香砂仁 | Zingiberaceae | Etlingera yunnanensis | 2 | VU | LC | 0.02 |
| 5 | 紫檀 | Leguminosae | Pterocarpus indicus | - | CR | EN | 0.07 |
| 6 | 毛叶樟 | Lauraceae | Camphora mollifolia | - | EN | EN | 0.03 |
| 7 | 云南石梓 | Lamiaceae | Gmelina arborea | - | VU | LC | 0.20 |
| 8 | 木橘 | Rutaceae | Aegle marmelos | - | VU | NT | 0.14 |
| 9 | 卡西松 | Pinaceae | Pinus kesiya | - | VU | LC | 0.06 |
| 10 | 辣椒砂仁 | Zingiberaceae | Amomum capsiciforme | - | - | CR | 0.08 |
| 11 | 厚叶沉香 | Thymelaeaceae | Aquilaria crassna | - | - | CR | 0.03 |
| 12 | 黄樟 | Lauraceae | Camphora parthenoxylon | - | - | EN | 0.06 |
| 13 | 喙花姜 | Zingiberaceae | Rhynchanthus beesianus | - | EN | - | 0.02 |
| 14 | 毛大叶臭花椒 | Rutaceae | Zanthoxylum myriacanthum var. pubescens | - | VU | - | 0.12 |
| 15 | 毛姜花 | Zingiberaceae | Hedychium villosum | - | VU | - | 0.03 |
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