Biodiv Sci ›› 2009, Vol. 17 ›› Issue (6): 533-548. DOI: 10.3724/SP.J.1003.2009.09253
Special Issue: 群落中的物种多样性:格局与机制
• Articles • Previous Articles Next Articles
Jingyun Fang1,*(), Xiangping Wang1,2, Zehao Shen1, Zhiyao Tang1, Jinsheng He1, Dan Yu3, Yuan Jiang4, Zhiheng Wang1, Chengyang Zheng1, Jiangling Zhu1, Zhaodi Guo1
Received:
2009-10-29
Accepted:
2009-11-27
Online:
2009-11-20
Published:
2009-11-20
Contact:
Jingyun Fang
Jingyun Fang, Xiangping Wang, Zehao Shen, Zhiyao Tang, Jinsheng He, Dan Yu, Yuan Jiang, Zhiheng Wang, Chengyang Zheng, Jiangling Zhu, Zhaodi Guo. Methods and protocols for plant community inventory[J]. Biodiv Sci, 2009, 17(6): 533-548.
分级 Scale | + | 1 | 2 | 3 | 4 | 5 |
---|---|---|---|---|---|---|
盖度范围 Range of coverage (%) | 少有出现 Rare | 0-5 | 5-25 | 25-50 | 50-75 | >75 |
Table 1 Braun-Blanquet scales of species coverage in plant communities
分级 Scale | + | 1 | 2 | 3 | 4 | 5 |
---|---|---|---|---|---|---|
盖度范围 Range of coverage (%) | 少有出现 Rare | 0-5 | 5-25 | 25-50 | 50-75 | >75 |
分级 Scale | 7 | 6 | 5 | 4 | 3 | 2 | 1 |
---|---|---|---|---|---|---|---|
符号 Symbol | soc | cop3 | cop2 | cop1 | sp | sol | un |
描述 Description | 极多 | 很多 | 多 | 尚多 | 不多 | 稀少 | 单株 |
Table 2 Drude scales of the species abundance in plant communities
分级 Scale | 7 | 6 | 5 | 4 | 3 | 2 | 1 |
---|---|---|---|---|---|---|---|
符号 Symbol | soc | cop3 | cop2 | cop1 | sp | sol | un |
描述 Description | 极多 | 很多 | 多 | 尚多 | 不多 | 稀少 | 单株 |
Fig. 2 Systematic sampling sites for plant community inventory in the North China and the adjacent areas. The background shows the distribution of vegetation based on the Vegetation Atlas of China (1:1,000,000) (Hou, 2001).
Fig. 3 Illustration of vertical structure of the forest comm- unity (from Lin, 1990). The plot is located at Erliping, Cuian County, Fujian, at an altitude of 770 m above sea level. The species are: 1, Castanopsis eyrei; 2, Cyclobalanopsis glauca; 3, Lithocarpus harlandii; 4, Photinia glabra; 5, Symplocos confusa; 6, Rhododendron latoucheae; 7, R. ovatum; 8, Lasianthus chinensis; 9, Oligostachyum oedogonatum; 10, Eurya nitida; 11, Hicriopteris chinensis.
样方编号 | 群落类型 | 样方面积 | ||||||||
---|---|---|---|---|---|---|---|---|---|---|
调查地点 | 省 县(林业局) 乡(林场) 村(林班) | |||||||||
具体位置描述: | ||||||||||
纬度 | 地形 | ( )山地 ( )洼地 ( )丘陵 ( )平原 ( )高原 | ||||||||
经度 | 坡位 | ( )谷地 ( )下部 ( )中下部 ( )中部 | ||||||||
海拔 | ( )中上部 ( )山顶 ( )山脊 | |||||||||
坡向 | 森林起源 | ( )原始林 ( )次生林 ( )人工林 | ||||||||
坡度 | 干扰程度 | ( )无干扰 ( )轻微 ( )中度 ( )强度 | ||||||||
土壤类型 | 林龄 | 群落剖面图: | ||||||||
垂直结构 | 层高(m) | 盖度(%) | 优势种 | |||||||
乔木层 | ||||||||||
亚乔木层 | ||||||||||
灌木层 | ||||||||||
草本层 | ||||||||||
调查人 | ||||||||||
记录人 | 调查日期 |
Appendix 1 Records for forest plots
样方编号 | 群落类型 | 样方面积 | ||||||||
---|---|---|---|---|---|---|---|---|---|---|
调查地点 | 省 县(林业局) 乡(林场) 村(林班) | |||||||||
具体位置描述: | ||||||||||
纬度 | 地形 | ( )山地 ( )洼地 ( )丘陵 ( )平原 ( )高原 | ||||||||
经度 | 坡位 | ( )谷地 ( )下部 ( )中下部 ( )中部 | ||||||||
海拔 | ( )中上部 ( )山顶 ( )山脊 | |||||||||
坡向 | 森林起源 | ( )原始林 ( )次生林 ( )人工林 | ||||||||
坡度 | 干扰程度 | ( )无干扰 ( )轻微 ( )中度 ( )强度 | ||||||||
土壤类型 | 林龄 | 群落剖面图: | ||||||||
垂直结构 | 层高(m) | 盖度(%) | 优势种 | |||||||
乔木层 | ||||||||||
亚乔木层 | ||||||||||
灌木层 | ||||||||||
草本层 | ||||||||||
调查人 | ||||||||||
记录人 | 调查日期 |
Fig. 4 Plot setting and quadrate coding for forest communities. The 20 m × 30 m plot is composed of six quadrates (A-F), each with an area of 10 m×10 m. The shadowed quadrates (S1 and S2) were selected for shrub layer investigation, and subplots (H1-5) were selected for herbaceous layer investigation. A buffering zone of 10-20 m at each side of the plot is necessary to keep the plot away from apparent human activities.
Fig. 7 Plot setting of shrub (grassland) communities. Five subplots (H1-5) were selected within each 10 m ×10 m plot. A buffering zone of 10 m at each side of the plot is necessary to keep the plot away from apparent human activities. For shrubs, whole plot (10 m × 10 m) should be investigated, and for grasslands, five subplots will be investigated.
属性 Traits | 建议的计量单位Unit suggested | 范围 Range | 最小重复数 Minimum replications | 推荐的重复数 Suggested replications | |
---|---|---|---|---|---|
叶片属性 Leaf | |||||
叶片大小 Leaf size | mm2 | 1-106 | 5, 2 | 10, 2 | |
叶片厚度 Leaf thickness | mm | 5, 2 | 10, 2 | ||
叶片干物质含量 Leaf dry matter content | mg/g | 50-700 | 5, 2 | 10, 2 | |
比叶面积 Specific leaf area | mm2/mg | 2-80 | 5, 2 | 10, 2 | |
叶寿命 Leaf lifespan | month | 0.5-200 | 3, 12 | 10, 12 | |
叶片N Leaf N concentration | mg/g | 10-60 | 5, 2 | 10, 2 | |
叶片P Leaf P concentration | mg/g | 0.5-5 | 5, 2 | 10, 2 | |
最大光合速率 Maximum photosynthetic rate | μmol·m-2·s-1 | 5, 2 | 10, 2 | ||
气孔导度 Leaf stomatal water conductance | mol·m-2·s-1 | 5, 2 | 10, 2 | ||
蒸腾速率 Leaf transpiration | mol·m-2·s-1 | 5, 2 | 10, 2 | ||
枝干属性 Stem | |||||
树皮厚度 Bark thickness | mm | 5 | 10 | ||
树干密度 Stem specific density | mg/mm3 | 0.4-1.2 | 5 | 10 | |
心材/边材比率 Heartwood to sapwood area ratio | unitless | 5 | 10 | ||
树干高度 Stem height | m | ? -100 | 10 | 25 | |
根系属性 Root | |||||
类型 Root type | cat. | 5 | 10 | ||
比根长 Specific root length | m/g | 10-500 | 5, 10 | 10, 10 | |
细根直径 Fine root diameter | mm | 5, 10 | 10, 10 | ||
组织密度 Tissue density | mg/mm3 | 5, 10 | 10, 10 | ||
细根N Fine root N concentration | mg/g | 5, 2 | 10, 2 | ||
细根P Fine root P concentration | mg/g | 5, 2 | 10, 2 | ||
繁殖属性 Reproduction | |||||
种子大小 Seed mass | mg | 10-3-107 | 3, 5 | 10, 5 | |
传播类型 Dispersal mode | cat. | 3 | 3 | ||
萌枝能力 Resprouting capacity | unitless | 0-100 | 5 | 25 |
Table 3 Metric units, range, minimum and suggested replications for plant trait measurements. The first and second numbers in the minimum (and suggested) replications represent replications of individuals and organs, respectively. This table is summarized from Cornelissen et al. (2003a), Pregitzer et al. (2002), and He et al. (2006b)
属性 Traits | 建议的计量单位Unit suggested | 范围 Range | 最小重复数 Minimum replications | 推荐的重复数 Suggested replications | |
---|---|---|---|---|---|
叶片属性 Leaf | |||||
叶片大小 Leaf size | mm2 | 1-106 | 5, 2 | 10, 2 | |
叶片厚度 Leaf thickness | mm | 5, 2 | 10, 2 | ||
叶片干物质含量 Leaf dry matter content | mg/g | 50-700 | 5, 2 | 10, 2 | |
比叶面积 Specific leaf area | mm2/mg | 2-80 | 5, 2 | 10, 2 | |
叶寿命 Leaf lifespan | month | 0.5-200 | 3, 12 | 10, 12 | |
叶片N Leaf N concentration | mg/g | 10-60 | 5, 2 | 10, 2 | |
叶片P Leaf P concentration | mg/g | 0.5-5 | 5, 2 | 10, 2 | |
最大光合速率 Maximum photosynthetic rate | μmol·m-2·s-1 | 5, 2 | 10, 2 | ||
气孔导度 Leaf stomatal water conductance | mol·m-2·s-1 | 5, 2 | 10, 2 | ||
蒸腾速率 Leaf transpiration | mol·m-2·s-1 | 5, 2 | 10, 2 | ||
枝干属性 Stem | |||||
树皮厚度 Bark thickness | mm | 5 | 10 | ||
树干密度 Stem specific density | mg/mm3 | 0.4-1.2 | 5 | 10 | |
心材/边材比率 Heartwood to sapwood area ratio | unitless | 5 | 10 | ||
树干高度 Stem height | m | ? -100 | 10 | 25 | |
根系属性 Root | |||||
类型 Root type | cat. | 5 | 10 | ||
比根长 Specific root length | m/g | 10-500 | 5, 10 | 10, 10 | |
细根直径 Fine root diameter | mm | 5, 10 | 10, 10 | ||
组织密度 Tissue density | mg/mm3 | 5, 10 | 10, 10 | ||
细根N Fine root N concentration | mg/g | 5, 2 | 10, 2 | ||
细根P Fine root P concentration | mg/g | 5, 2 | 10, 2 | ||
繁殖属性 Reproduction | |||||
种子大小 Seed mass | mg | 10-3-107 | 3, 5 | 10, 5 | |
传播类型 Dispersal mode | cat. | 3 | 3 | ||
萌枝能力 Resprouting capacity | unitless | 0-100 | 5 | 25 |
属性 Traits | 测定方法 Measuring methods |
---|---|
叶片属性 Leaf 叶片大小 Leaf size | 叶面积仪(如Delta-T (Cambridge, UK), Li-Cor (Lincoln, Nebraska, USA)), 或扫描仪 |
叶片厚度 Leaf thickness | 显微镜或游标卡尺测量 |
叶片干物质含量 Leaf dry matter content | 叶片装入自封袋, 加水浸湿, 保持12 h以上, 吸掉多余水分, 称鲜重; 60℃烘干至少72 h(或80℃至少48 h), 称干重, 计算 |
比叶面积 Specific leaf area | 面积测定: 叶面积仪(如Delta-T (Cambridge, UK), Li-Cor (Lincoln, Nebraska, USA)。干重测定: 烘箱60℃烘烤至少72 h(或80℃至少48 h), 称干重。由叶面积和干重计算比叶面积。 |
叶寿命 Leaf lifespan | 双子叶植物: 叶片标记法; 单子叶植物: 测定叶片特定区域的生产和死亡动态, 具体见 |
叶片N Leaf N concentration | 元素分析仪法(2400Ⅱ CHNS/O Elemental Analyzer, Perkin-Elmer, Boston, MA,USA)( |
叶片P Leaf P concentration | 钼蓝比色法 ( |
最大光合速率 Pnmax | Li-6400便携式光合仪测定, 建议设光照=1,500 μmol/(m2·s1), [CO2] = 380 ppm |
气孔导度 Gs | Li-6400便携式光合仪测定 |
蒸腾速率 Tr | Li-6400便携式光合仪测定 |
枝干属性 Stem | |
树皮厚度 Bark thickness | 游标卡尺测量 |
树干密度 Stem specific density | 体积替代法: 鲜树干完全浸于盛水的量筒中约5 s, 读取增加的体积; 烘箱中60℃烘烤至少72 h (小树干)或96 h (大树干), 称干重。由体积和重量计算树干密度 |
心材/边材比率 Heartwood to sapwood area ratio | 钻取生长芯, 经打磨处理后, 用高精度扫描仪进行图像扫描, 据心材和边材颜色区分其界限, 用年轮分析软件测量每个生长芯的树皮厚度、边材宽度和边材年轮数。以树干横断面为标准圆形计算心材/边材比率 |
树干高度 Stem height | 直接测量法(如激光测距仪)和三角函数法 |
根系属性 Root | |
类型 Root type | 参照 |
比根长 Specific root length | 解剖镜下区分出活的、完整的根, 用目镜测微尺测量长度; 或用WinRHIZO根系分析系统, 60℃下至少烘烤72 h(或80℃至少48 h), 称重, 然后计算比根长 |
细根直径 Fine root diameter | 解剖镜下区分出活的、完整的根, 用目镜测微尺校正细根直径 |
组织密度 Tissue density | 根干重/根体积, 详细内容参见 |
细根N Fine root N concentration | 元素分析仪法(如CHNS/O Elemental Analyzer, Perkin-Elmer, Boston, MA, USA)或微量凯氏定氮法 |
细根P Fine root N concentration | 钼蓝比色法 ( |
繁殖属性 Reproduction | |
种子大小 Seed mass | 野外采成熟果实, 清理(洗)出种子, 统计种子数目, 80℃下至少烘烤48 h, 直到恒重, 称重。 |
传播类型 Dispersal mode | 记录所有与传播有关的分类学特征, 据重要性按降序排列。具体分类特征见 |
萌枝能力 Resprouting capacity | 根据萌枝数, 划分等级 |
Table 4 Measuring methods for ecological traits of plants
属性 Traits | 测定方法 Measuring methods |
---|---|
叶片属性 Leaf 叶片大小 Leaf size | 叶面积仪(如Delta-T (Cambridge, UK), Li-Cor (Lincoln, Nebraska, USA)), 或扫描仪 |
叶片厚度 Leaf thickness | 显微镜或游标卡尺测量 |
叶片干物质含量 Leaf dry matter content | 叶片装入自封袋, 加水浸湿, 保持12 h以上, 吸掉多余水分, 称鲜重; 60℃烘干至少72 h(或80℃至少48 h), 称干重, 计算 |
比叶面积 Specific leaf area | 面积测定: 叶面积仪(如Delta-T (Cambridge, UK), Li-Cor (Lincoln, Nebraska, USA)。干重测定: 烘箱60℃烘烤至少72 h(或80℃至少48 h), 称干重。由叶面积和干重计算比叶面积。 |
叶寿命 Leaf lifespan | 双子叶植物: 叶片标记法; 单子叶植物: 测定叶片特定区域的生产和死亡动态, 具体见 |
叶片N Leaf N concentration | 元素分析仪法(2400Ⅱ CHNS/O Elemental Analyzer, Perkin-Elmer, Boston, MA,USA)( |
叶片P Leaf P concentration | 钼蓝比色法 ( |
最大光合速率 Pnmax | Li-6400便携式光合仪测定, 建议设光照=1,500 μmol/(m2·s1), [CO2] = 380 ppm |
气孔导度 Gs | Li-6400便携式光合仪测定 |
蒸腾速率 Tr | Li-6400便携式光合仪测定 |
枝干属性 Stem | |
树皮厚度 Bark thickness | 游标卡尺测量 |
树干密度 Stem specific density | 体积替代法: 鲜树干完全浸于盛水的量筒中约5 s, 读取增加的体积; 烘箱中60℃烘烤至少72 h (小树干)或96 h (大树干), 称干重。由体积和重量计算树干密度 |
心材/边材比率 Heartwood to sapwood area ratio | 钻取生长芯, 经打磨处理后, 用高精度扫描仪进行图像扫描, 据心材和边材颜色区分其界限, 用年轮分析软件测量每个生长芯的树皮厚度、边材宽度和边材年轮数。以树干横断面为标准圆形计算心材/边材比率 |
树干高度 Stem height | 直接测量法(如激光测距仪)和三角函数法 |
根系属性 Root | |
类型 Root type | 参照 |
比根长 Specific root length | 解剖镜下区分出活的、完整的根, 用目镜测微尺测量长度; 或用WinRHIZO根系分析系统, 60℃下至少烘烤72 h(或80℃至少48 h), 称重, 然后计算比根长 |
细根直径 Fine root diameter | 解剖镜下区分出活的、完整的根, 用目镜测微尺校正细根直径 |
组织密度 Tissue density | 根干重/根体积, 详细内容参见 |
细根N Fine root N concentration | 元素分析仪法(如CHNS/O Elemental Analyzer, Perkin-Elmer, Boston, MA, USA)或微量凯氏定氮法 |
细根P Fine root N concentration | 钼蓝比色法 ( |
繁殖属性 Reproduction | |
种子大小 Seed mass | 野外采成熟果实, 清理(洗)出种子, 统计种子数目, 80℃下至少烘烤48 h, 直到恒重, 称重。 |
传播类型 Dispersal mode | 记录所有与传播有关的分类学特征, 据重要性按降序排列。具体分类特征见 |
萌枝能力 Resprouting capacity | 根据萌枝数, 划分等级 |
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