生物多样性 ›› 2019, Vol. 27 ›› Issue (2): 168-176. DOI: 10.17520/biods.2018224
涂伟凤1,张洋2,3,*(),汤洁2,3,涂玉琴2,3,辛佳佳2,3,姬红利4,张南峰2,张弢2
收稿日期:
2018-08-13
接受日期:
2019-01-02
出版日期:
2019-02-20
发布日期:
2019-04-16
通讯作者:
张洋
基金资助:
Tu Weifeng1,Zhang Yang2,3,*(),Tang Jie2,3,Tu Yuqin2,3,Xin Jiajia2,3,Ji Hongli4,Zhang Nanfeng2,Zhang Tao2
Received:
2018-08-13
Accepted:
2019-01-02
Online:
2019-02-20
Published:
2019-04-16
Contact:
Zhang Yang
摘要:
印度蔊菜(Rorippa indica)与无瓣蔊菜(R. dubia)的分类关系仍存在较大争议, 为阐明二者的分类学关系, 本研究综合利用形态指标测量、DNA相对含量检测、体细胞染色体观察和基于SSR分子标记的遗传距离分析等方法, 系统地比较了二者的分类学特征和细胞遗传学差异。结果表明: 印度蔊菜(2n = 48)为六倍体, 无瓣蔊菜(2n = 32)为四倍体。同时, 前者DNA相对含量约为150, 是后者的1.5倍。通过45对SSR分子标记的遗传距离分析得出, 相对于球果蔊菜(R. globosa)与欧亚蔊菜(R. sylvestris), 二者亲缘关系较近, 独立聚类为一支, 但在遗传距离为0.23处可以明确划分为两个种。同时, 利用角果长度和结籽密度两个形态指标可以将二者明显区分为两个种。另外, 二者存在明显的生殖障碍, 通过正反交授粉实验得出: 印度蔊菜与无瓣蔊菜自花授粉的结实率分别为97.73%和95.65%, 而以印度蔊菜为母本的杂交处理结实率为0, 以无瓣蔊菜为母本的杂交处理结实率为47.06%, 但其种子萌发率为0。综上所述, 印度蔊菜与无瓣蔊菜为两个种。
涂伟凤,张洋,汤洁,涂玉琴,辛佳佳,姬红利,张南峰,张弢 (2019) 印度蔊菜与无瓣蔊菜形态变异特征的比较及分类关系. 生物多样性, 27, 168-176. DOI: 10.17520/biods.2018224.
Tu Weifeng,Zhang Yang,Tang Jie,Tu Yuqin,Xin Jiajia,Ji Hongli,Zhang Nanfeng,Zhang Tao (2019) Comparison of taxonomic morphological characteristics between Rorippa indica and R. dubia. Biodiversity Science, 27, 168-176. DOI: 10.17520/biods.2018224.
居群代号 Population code | 地点 Locality | 生境 Habitat | 经纬度 Location | 海拔 Altitude (m) |
---|---|---|---|---|
印度蔊菜居群 Rorippa indica | ||||
Ri-1 | 江西省大余县池江镇 Chijiang Town, Dayu, Jiangxi | 田埂和排水渠两侧草丛附近 Near the balk and offtake of the brushwood | 25.49° N; 114.55° E | 174 |
Ri-2 | 江西省大余县河洞乡 Hedong Town, Dayu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 25.31° N; 114.08° E | 372 |
Ri-3 | 江西省永修县大屋村 Dawu Village, Yongxiu, Jiangxi | 溪流旁的碎石滩周围 Near the rocky beach of streams | 29.17° N; 115.57° E | 32 |
Ri-4 | 江西省安福县岭上村 Lingshang Village, Anfu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 27.20° N; 114.58° E | 89 |
Ri-5 | 江西省永修县内江村 Neijiang Village, Yongxiu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 29.04° N; 115.81o E | 56 |
Ri-6 | 江西省农业科学院 Jiangxi Academy of Agricultural Sciences, Nanchang, Jiangxi | 有房屋遮阴, 生于排水沟周围 Around the drain and under the eaves | 28.56° N; 115.94o E | 45 |
无瓣蔊菜居群 R. dubia | ||||
Rd-1 | 江西省永修县陈司村 Chensi Village, Yongxiu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 29.31° N; 115.44° E | 79 |
Rd-2 | 江西省永修县南坑村 Nankeng Village, Yongxiu, Jiangxi | 有树木遮阴, 路旁排水沟两侧 Around the drain and under the trees | 29.14° N; 115.53° E | 203 |
Rd-3 | 江西省大余县樟斗镇 Zhangdou Town, Dayu, Jiangxi | 冬闲的水稻田内 In the winter leisure farmland | 25.58° N; 114.47° E | 296 |
Rd-4 | 江西省永修县河桥村 Heqiao Village, Yongxiu, Jiangxi | 农家菜园内, 土壤肥沃 In the garden with fertile soil | 29.29° N; 115.56° E | 69 |
球果蔊菜居群 R. globosa | ||||
Rg-1 | 江西省永修县吴城村 Wucheng Village, Yongxiu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 29.18° N; 116.00° E | 28 |
Rg-2 | 江西省永修县丁山村 Dingshan Village, Yongxiu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 29.14° N; 115.98° E | 33 |
欧亚蔊菜居群 R. sylvestris | ||||
Rs-1 | 江西省西宁市城北区 Chengbei District, Xining, Qinghai | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 36.73° N; 101.75° E | 2,206 |
Rs-2 | 江西省西宁市城北区 Chengbei District, Xining, Qinghai | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 36.76° N; 101.73° E | 2,268 |
表1 本研究中印度蔊菜、无瓣蔊菜、球果蔊菜和欧亚蔊菜居群的生境与地理信息
Table 1 Habitat and geographic information of Rorippa indica, R. dubia, R. globosa and R. sylvestris
居群代号 Population code | 地点 Locality | 生境 Habitat | 经纬度 Location | 海拔 Altitude (m) |
---|---|---|---|---|
印度蔊菜居群 Rorippa indica | ||||
Ri-1 | 江西省大余县池江镇 Chijiang Town, Dayu, Jiangxi | 田埂和排水渠两侧草丛附近 Near the balk and offtake of the brushwood | 25.49° N; 114.55° E | 174 |
Ri-2 | 江西省大余县河洞乡 Hedong Town, Dayu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 25.31° N; 114.08° E | 372 |
Ri-3 | 江西省永修县大屋村 Dawu Village, Yongxiu, Jiangxi | 溪流旁的碎石滩周围 Near the rocky beach of streams | 29.17° N; 115.57° E | 32 |
Ri-4 | 江西省安福县岭上村 Lingshang Village, Anfu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 27.20° N; 114.58° E | 89 |
Ri-5 | 江西省永修县内江村 Neijiang Village, Yongxiu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 29.04° N; 115.81o E | 56 |
Ri-6 | 江西省农业科学院 Jiangxi Academy of Agricultural Sciences, Nanchang, Jiangxi | 有房屋遮阴, 生于排水沟周围 Around the drain and under the eaves | 28.56° N; 115.94o E | 45 |
无瓣蔊菜居群 R. dubia | ||||
Rd-1 | 江西省永修县陈司村 Chensi Village, Yongxiu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 29.31° N; 115.44° E | 79 |
Rd-2 | 江西省永修县南坑村 Nankeng Village, Yongxiu, Jiangxi | 有树木遮阴, 路旁排水沟两侧 Around the drain and under the trees | 29.14° N; 115.53° E | 203 |
Rd-3 | 江西省大余县樟斗镇 Zhangdou Town, Dayu, Jiangxi | 冬闲的水稻田内 In the winter leisure farmland | 25.58° N; 114.47° E | 296 |
Rd-4 | 江西省永修县河桥村 Heqiao Village, Yongxiu, Jiangxi | 农家菜园内, 土壤肥沃 In the garden with fertile soil | 29.29° N; 115.56° E | 69 |
球果蔊菜居群 R. globosa | ||||
Rg-1 | 江西省永修县吴城村 Wucheng Village, Yongxiu, Jiangxi | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 29.18° N; 116.00° E | 28 |
Rg-2 | 江西省永修县丁山村 Dingshan Village, Yongxiu, Jiangxi | 废旧房屋墙角碎石缝隙中 Narrow gaps in the rubble near the corner of old houses | 29.14° N; 115.98° E | 33 |
欧亚蔊菜居群 R. sylvestris | ||||
Rs-1 | 江西省西宁市城北区 Chengbei District, Xining, Qinghai | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 36.73° N; 101.75° E | 2,206 |
Rs-2 | 江西省西宁市城北区 Chengbei District, Xining, Qinghai | 空旷的沙土地, 植被较少 Open sandy land with less vegetation | 36.76° N; 101.73° E | 2,268 |
图1 印度蔊菜与无瓣蔊菜叶片DNA相对含量。(a)印度蔊菜居群Ri-2; (b)无瓣蔊菜居群Rd-4; (c)印度蔊菜居群Ri-6; (d)无瓣蔊菜居群Rd-1; (e)印度蔊菜居群Ri-1; (f)无瓣蔊菜居群Rd-2; (g)印度蔊菜居群Ri-3; (h)无瓣蔊菜居群Rd-3。居群代号同表1。
Fig. 1 Relative content of DNA in leaves of Rorippa indica and R. dubia. (a) R. indica Ri-2; (b) R. dubia Rd-4; (c) R. indica Ri-6; (d) R. dubia Rd-1; (e) R. indica Ri-1; (f) R. dubia Rd-2; (g) R. indica Ri-3; h, R. dubia Rd-3. Population codes see Table 1.
图2 印度蔊菜与无瓣蔊菜子房壁细胞染色体中期分裂相观察。(a)印度蔊菜居群Ri-2; (b)无瓣蔊菜居群Rd-4; (c)印度蔊菜居群Ri-6; (d)无瓣蔊菜居群Rd-1; (e)印度蔊菜居群Ri-1; (f)瓣蔊菜居群Rd-2; (g)印度蔊菜居群Ri-3; (h)无瓣蔊菜居群Rd-3。居群代号同表1。
Fig. 2 Observation of metaphase division phase from the ovary parietal cells of Rorippa indica and Rorippa dubia. (a) R. indica Ri-2; (b) R. dubia Rd-4; (c) R. indica Ri-6; (d) R. dubia Rd-1; (e) R. indica Ri-1; (f) R. dubia Rd-2; (g) R. indica Ri-3; (h) R. dubia Rd-3. Population codes see Table 1.
图3 印度蔊菜、无瓣蔊菜、球果蔊菜和欧亚蔊菜居群的遗传距离。居群代号同表1。
Fig. 3 Genetic distance of Rorippa indica, R. dubia, and R. globosaand R. sylvestris. Population codes see Table 1.
处理 Treatment | 每花花瓣数 Petal number per flower | 种子列数 Seed column | 角果长度 Pod length (mm) | 每角果种子数 Number of seeds per pod | 结籽密度 (粒/cm) Seed density (No./cm) |
---|---|---|---|---|---|
印度蔊菜盆栽对照处理 Control for R. indica | 3.94 ± 0.24 d | 2 | 16.01 ± 1.17 b | 82.00 ± 7.20 c | 5.12 ± 0.15 d |
印度蔊菜盆栽干旱处理 Drought stress for R. indica | 3.06 ± 1.24 c | 2 | 10.19 ± 3.50 a | 43.92 ± 23.76 a | 4.07 ± 1.15 c |
印度蔊菜自然条件下生长 Natural condition for R. indica | 3.89 ± 0.32 d | 2 | 16.98 ± 1.06 b | 93.70 ± 3.92 c | 5.45 ± 0.33 d |
无瓣蔊菜盆栽对照处理 Control for R. dubia | 0.43 ± 0.84 a | 1 | 29.51 ± 5.21 d | 65.15 ± 16.13 b | 2.20 ± 0.31 a |
无瓣蔊菜盆栽干旱处理 Drought stress for R. dubia | 1.35 ± 1.11 b | 1 | 20.20 ± 2.87 c | 68.71 ± 10.38 b | 3.41 ± 0.38 b |
无瓣蔊菜自然条件下生长 Natural condition for R. dubia | 0.75 ± 1.20 a | 1 | 27.30 ± 5.24 d | 66.17 ± 18.92 b | 2.39 ± 0.38 a |
表2 干旱处理对印度蔊菜与无瓣蔊菜植株形态指标的影响
Table 2 Effects of drought treatments on plant morphology of Rorippa indica and R. dubia
处理 Treatment | 每花花瓣数 Petal number per flower | 种子列数 Seed column | 角果长度 Pod length (mm) | 每角果种子数 Number of seeds per pod | 结籽密度 (粒/cm) Seed density (No./cm) |
---|---|---|---|---|---|
印度蔊菜盆栽对照处理 Control for R. indica | 3.94 ± 0.24 d | 2 | 16.01 ± 1.17 b | 82.00 ± 7.20 c | 5.12 ± 0.15 d |
印度蔊菜盆栽干旱处理 Drought stress for R. indica | 3.06 ± 1.24 c | 2 | 10.19 ± 3.50 a | 43.92 ± 23.76 a | 4.07 ± 1.15 c |
印度蔊菜自然条件下生长 Natural condition for R. indica | 3.89 ± 0.32 d | 2 | 16.98 ± 1.06 b | 93.70 ± 3.92 c | 5.45 ± 0.33 d |
无瓣蔊菜盆栽对照处理 Control for R. dubia | 0.43 ± 0.84 a | 1 | 29.51 ± 5.21 d | 65.15 ± 16.13 b | 2.20 ± 0.31 a |
无瓣蔊菜盆栽干旱处理 Drought stress for R. dubia | 1.35 ± 1.11 b | 1 | 20.20 ± 2.87 c | 68.71 ± 10.38 b | 3.41 ± 0.38 b |
无瓣蔊菜自然条件下生长 Natural condition for R. dubia | 0.75 ± 1.20 a | 1 | 27.30 ± 5.24 d | 66.17 ± 18.92 b | 2.39 ± 0.38 a |
处理 Treatment | 授粉花朵数 No. of pollinated flowers | 结实角果数 No. of developed pods | 结实率 Seed setting rate (%) | 种子列数 Seed column | 角果长度 Pod length (mm) | 每角果 种子数 No. of seeds per pod | 种子萌发率 Seed germination rate (%) |
---|---|---|---|---|---|---|---|
印度蔊菜自花授粉 Self pollination of R. indica | 44 | 43 | 97.73 | 2 | 15.23 ± 2.16 a | 90.26 ± 3.89 a | 96.70 |
无瓣蔊菜自花授粉 Self pollination of R. dubia | 46 | 44 | 95.65 | 1 | 29.45 ± 6.52 b | 63.28 ± 10.67 b | 97.80 |
♀印度蔊菜 × ♂无瓣蔊菜 ♀R. indica × ♂R. dubia | 36 | 0 | 0 | - | - | - | - |
♀无瓣蔊菜 × ♂印度蔊菜 ♀R. dubia × ♂R. indica | 34 | 16 | 47.06 | 1 | 23.97 ± 8.83 b | 46.16 ± 15.37 b | 0 |
表3 印度蔊菜与无瓣蔊菜杂交授粉实验结果
Table 3 Cross pollination experiment of Rorippa indica and R. dubia
处理 Treatment | 授粉花朵数 No. of pollinated flowers | 结实角果数 No. of developed pods | 结实率 Seed setting rate (%) | 种子列数 Seed column | 角果长度 Pod length (mm) | 每角果 种子数 No. of seeds per pod | 种子萌发率 Seed germination rate (%) |
---|---|---|---|---|---|---|---|
印度蔊菜自花授粉 Self pollination of R. indica | 44 | 43 | 97.73 | 2 | 15.23 ± 2.16 a | 90.26 ± 3.89 a | 96.70 |
无瓣蔊菜自花授粉 Self pollination of R. dubia | 46 | 44 | 95.65 | 1 | 29.45 ± 6.52 b | 63.28 ± 10.67 b | 97.80 |
♀印度蔊菜 × ♂无瓣蔊菜 ♀R. indica × ♂R. dubia | 36 | 0 | 0 | - | - | - | - |
♀无瓣蔊菜 × ♂印度蔊菜 ♀R. dubia × ♂R. indica | 34 | 16 | 47.06 | 1 | 23.97 ± 8.83 b | 46.16 ± 15.37 b | 0 |
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