Following (1) the large-scale molecular phylogeny of seed plants based on plastid rbcL gene sequences (published in 1993 by Chase et al., Ann. Missouri Bot. Gard. 80:528-580) and (2) the 18S nuclear phylogeny of flowering plants (published in 1997 by Soltis et al., Ann. Missouri Bot. Gard. 84:1-49), we present a phylogenetic analysis of flowering plants based on a second plastid gene, atpB, analyzed separately and in combination with rbcL sequences for 357 taxa. Despite some discrepancies, the atpB-based phylogenetic trees were highly congruent with those derived from the analysis of rbcL and 18S rDNA, and the combination of atpB and rbcL DNA sequences (comprising approximately 3000 base pairs) produced increased bootstrap support for many major sets of taxa. The angiosperms are divided into two major groups: noneudicots with inaperturate or uniaperturate pollen (monocots plus Laurales, Magnoliales, Piperales, Ceratophyllales, and Amborellaceae-Nymphaeaceae-Illiciaceae) and the eudicots with triaperturate pollen (particularly asterids and rosids). Based on rbcL alone and atpB/rbcL combined, the noneudicots (excluding Ceratophyllum) are monophyletic, whereas in the atpB trees they form a grade. Ceratophyllum is sister to the rest of angiosperms with rbcL alone and in the combined atpB/rbcL analysis, whereas with atpB alone, Amborellaceae, Nymphaeaceae, and Illiciaceae/Schisandraceae form a grade at the base of the angiosperms. The phylogenetic information at each codon position and the different types of substitutions (observed transitions and transversions in the trees vs. pairwise comparisons) were examined; taking into account their respective consistency and retention indices, we demonstrate that third-codon positions and transitions are the most useful characters in these phylogenetic reconstructions. This study further demonstrates that phylogenetic analysis of large matrices is feasible.

<p id="p00005"><strong>Background &#x00026; Aim:</strong> A high-quality checklist is of great significance to plant taxonomy, ecology, geography, and the utilization of plant resources. Unfortunately, since the 1980s, the checklist of Heilongjiang Province has not been fully updated for a long time, which has seriously hindered related research. This paper attempts to obtain a credible province-wide checklist and county-level plant distribution data. <br> <strong>Method:</strong> The TNRS (Taxonomic Name Resolution) was used to correct the spelling of scientific names. And the scientific name adopts the viewpoints in the Catalogue of Life China (2021). The checklist mainly refers to documents, i.e. Catalogue of Life China (2021), <i>Key of Plants of Northeastern China </i>(2nd edn),<i> Flora of Heilongjiang</i>,<i> Ligneous Flora of Heilongjiang</i>,<i> Distribution Atlas of Northeast China Plant</i>,<i> Flora Plantarum Herbacearum Chinae Boreali-Orientalis</i>, some articles and specimen data (from NSII, GBIF, NEFI, NEAU, IFP) to locate the species distribution in county-level divisions. <br> <strong>Results:</strong> A total of 2,276 species (subspecies and varieties) belonging to 651 genera and 132 families of tracheophyte in Heilongjiang Province were counted, including 17 species belonging to 6 genera and 2 families of lycopods, 81 species belonging to 34 genera, and 16 families of pteridophytes, 20 species belonging to 6 genera and 3 families of gymnosperms, and 2,158 species belonging to 605 genera and 111 families of angiosperms. Among them the families of Asteraceae (67 genera, 258 species), Poaceae (61 genera, 187 species), Cyperaceae (14 genera, 174 species), Ranunculaceae (18 genera, 124 species) and Rosaceae (24 genera, 112 species) contain a large number of species. This checklist shows remarkable differences in species distribution among counties in Heilongjiang Province. About 17 percent of 131 counties have more than 1,000 species, and the county-level distribution data are seriously unbalanced. <br> <strong>Conclusions:</strong> The provincial checklist is reliable, but the quality of county-level distribution data needs to be significantly improved.</p>

黑龙江省位于中国的东北端, 省下辖区划分复杂而繁多, 植被可分为3大区域: 寒温带针叶林区域、温带针阔混交林区域、温带草原区域。本名录主要参考《中国生物物种名录(2021版)》《东北植物志》《东北植物检索表》《黑龙江省植物志》《黑龙江省树木志》《东北植物分布图集》《东北草本植物志》等著作、近年来发表的论文和来自国家标本资源共享平台（NSII）、全球生物多样性信息网络(GBIF)、东北林业大学植物标本室(NEFI)、东北农业大学生命科学学院植物标本室(NEAU)、中国科学院沈阳应用生态研究所东北生物标本馆(IFP)的标本资料以及最近发表的文献资料。由于&#x0201c;无分布&#x0201d;比&#x0201c;有分布&#x0201d;的信息相对较难确定, 我们在整理数据时, 轻易不删除重要著作中已经有收录的物种, 除非有较可信的证据, 这可能使得本数据集的物种数量稍偏多。本名录中物种分布精确到县级, 共收录黑龙江省野生维管植物132科651属2,276种(亚种、变种), 有凭证标本的本土植物2,122种(亚种、变种), 广泛入侵的44种, 无凭证标本但是有较可靠资料的154种。其中石松类2科6属17种, 蕨类植物16科34属81种, 裸子植物3科6属20种, 被子植物111科605属2,158种。收录国家级重点保护植物22科25属39种。黑龙江野生维管植物中, 菊科(67属258种)、禾本科(61属187种)、莎草科(14属174种)、毛茛科(18属124种)和蔷薇科(24属112种)所含种数较多。统计发现, 黑龙江省物种县域分布差异较大, 131个县级行政单位中, 物种数超过1,000的约占1/6, 且县级分布数据严重不均衡。本名录中, 省级名录的可信度相对较高, 县级分布数据质量还需要大大提升。

通过文献检索，统计分析2015～2017年间文献报道的福建省境内新发现的维管束植物，包括新种11种(隶属5科)和新记录种80种(隶属35科)。至此，福建维管束植物达252科1643属5064种。

Background: Lycophytes and ferns are the second largest group of vascular plants, dating back as far as 400 million years ago. Before the domination of angiosperms, lycophytes and ferns were the key component of ancient terrestrial ecosystems, and are still important to modern ecosystems today. Since the 1940s, the study of lycophytes and ferns in China has attracted worldwide attention. In the past five years since the 19th International Botanical Congress (IBC) held in Shenzhen in 2017, research on lycophytes and ferns in China has expanded internationally and focuses on a wider range of scientific issues. A series of important advances have been made on plant diversity, conservation, phylogeny, and ecological adaptation. Results: From 2017 to 2022, the tree of life for lycophytes and ferns in the world has been reconstructed by the family level, and a new evolutionary model for the sporangium was proposed. Some of the phylogenetic and taxonomic problems of order, family, genus and species of lycophytes and ferns were solved, meanwhile 106 new taxa were published. A large number of floristic collections and studies have been conducted, and six Chinese plus one worldwide monographs on the diversity of lycophytes and ferns have been published. In addition, 65 species of lycophytes and ferns under state priority conservation have been ex situ protected, and spore reproduction has been achieved in priority conservation groups such as Cyatheaceae, Ceratopteris, Angiopteris and Platycerium wallichiia. The ecological restoration and adaptive evolution of lycophytes and ferns were studied under the phylogeny. Perspectives: After literature survey on the research of lycophytes and ferns in China from 2017 to 2022, we make the following suggestions for future studies: (1) Focus future research on Chinese oligotypic families/genera and the worldwide mega-diverse families/genera. (2) Prioritize research in under-studied areas, such as Tibet and Sichuan, and utilize new technologies (e.g., DNA barcoding) to improve the efficiency and accuracy of species identification. (3) When clarifying the systematic relationship between families, genera, and species by using multi-disciplinary approaches, it is necessary to conduct research on plant evolutionary history and adaptation mechanisms. (4) Research efforts should focus on the common scientific conjecture of the systematic position of lycophytes and fern as the origin for the evolution of terrestrial vascular plants. (5) Pay more attention to the cooperation among the phylogenetics, ecology, phytochemistry, conservation biology and other subjects of lycophytes and ferns.

石松类和蕨类植物是维管植物的第二大类群, 其起源可追溯到4亿年前。在被子植物出现之前, 石松类和蕨类植物在古地球生态系统中占主导地位, 其重要性一直延续到现在。自20世纪40年代开始, 中国石松类和蕨类植物研究就令世界瞩目, 尤其是2017年第19届国际植物学大会在中国深圳召开后的5年时间里, 中国石松类和蕨类植物研究更是面向世界、走向国际, 研究更为广泛的科学问题, 在物种多样性、保护、系统演化和生态适应性等方面取得了一系列重要研究进展。2017-2022年, 多个中国研究团队利用多组学数据构建了世界石松类和蕨类植物科级水平的生命之树并提出了关键性状孢子囊环带演化的新模式; 解决了石松类和蕨类植物中目级、科级、属级和种级众多关键的系统分类学等问题, 发表了106个新分类群; 开展了大量的植物区系调查和研究, 出版了6部中国石松类和蕨类植物多样性专著和1部世界性专著; 对65种国家重点保护的石松类和蕨类植物进行了迁地保护, 同时实现了桫椤科、水蕨属(Ceratopteris)、观音座莲属(Angiopteris)和鹿角蕨(Platycerium wallichii)等重点保护类群的孢子繁殖; 在系统发育框架下, 研究了石松类和蕨类植物的生态修复功能和生态适应性演化。通过对2017-2022年研究成果的总结和思考, 本文对未来石松类和蕨类植物的发展提出以下建议: (1)提高中国寡型科属以及世界性大科大属的关注力度; (2)加强西藏、四川等薄弱地区石松类和蕨类植物的调查研究, 并结合新技术, 如DNA条形码等以提高区系调查中物种鉴别的效率和准确性; (3)运用多学科交叉的研究方法厘清各科、属、种间系统关系的同时, 还应加强系统和生态适应性演化之间的协同研究; (4)关注石松类和蕨类植物系统位置作为陆生维管植物演化起点的共性科学问题; (5)加强石松类和蕨类植物系统分类学与生态学、植物化学、保护生物学等学科间交叉合作研究。

<p id="p00005"><strong>Background:</strong> Naturalization is the pre-invasion stage. Detailed research on naturalized plants is a fundamental first step for scientific management of alien species. This research is necessary for predicting and hence preventing such invasion caused by alien plants. Regional studies of naturalized plants are important for local governments to manage and control of such plants. As an important area for foreign exchanges, East China is facing pressures from imported alien plants being naturalized and then invading. This study aims to enrich the basic information about naturalized plants of East China, discussed the current deficiencies in the study of naturalized plants, and summarized the direction of future research. </p> <p id="p00010"><strong>Methods:</strong> We conducted a comprehensive survey of naturalized plants in East China (Anhui, Fujian, Jiangsu, Jiangxi, Shandong, Shanghai and Zhejiang provinces) from 2014 to 2019 using literature reports, specimen information, and taxonomic research. We analyzed the species composition and distribution pattern of naturalized plants in East China and conducted a comprehensive analysis of their first detected locations, introduced route, and the time when species were discovered. </p> <p id="p00015"><strong>Results:</strong> We found that 299 species belonging to 181 genera and 62 families were naturalized in East China. The four most dominant families were Compositae (49 species), Leguminosae (35 species), Gramineae (28 species) and Amaranthaceae (25 species). The four most dominant genera were <i>Amaranthus</i> (16 species), <i>Euphorbia</i>(10 species),<i>Ipomoea</i> (9 species) and<i>Solanum</i> (9 species). There was a large diversity in species identified as naturalized, with a high proportion of plants coming from the Americas, and concentration in introduction routes. The Americas are the primary geographical origin of naturalized alien plant species, with more species originating from North America rather than South America. The route of introduction analysis revealed that 143 naturalized plant species were introduced unintentionally due to human activity and 156 species were introduced intentionally. Ninety-five frequencies were introduced as ornamental plants, which take up 55.2% of all the frequencies introduced intentionally. This suggests that introductions of plants for the purpose of ornamental use is the most common way for species naturalization to occur. A comparative study of the first detected locations show that the largest number of naturalized species were firstly introduced to South China (84 species) which mostly went to the Guangdong Province (33 species) and Hong Kong District (33 species). This was followed by East China (74 species), the Taiwan Province (72 species), Southwest China (19 species, mainly from Yunnan), Northeast China (15 species, mainly from Liaoning), Northwest China (14 species, mainly from Xinjiang) and North China (14 species, mainly from Beijing). There were only two naturalized species found in Central China. Jiangsu, Fujian and Zhejiang provinces were the main importing areas for East China. This may be due to many factors such as international communication, population density, climate similarity, disturbance levels, biological characteristics unique to species and investigation degree. Spatially, the species diversity of the Fujian Province (236 species) is significantly higher than that of other provinces, but little difference has been detected among other provinces. There are 57 species that are only distributed in Fujian (19.1%), which is the most when compared with other provinces. The amount of foreign exchange and climate similarity between native habitat and naturalized place may be the primary driver of current diversity and distribution pattern. Historically, the naturalized plants in East China showed an exponential growth trend after 1850. The growth rate is 1.5 species per year, and is currently in a period of rapid growth. In the 21st century, more than 80% of naturalized plants have been unintentionally introduced. Therefore, special attention should be placed on preventing the introduction of unintentional species. </p> <p id="p00020"><strong>Proposal:</strong> Constructing an effective risk assessment system of alien plants based on an accurate and complete database of naturalized plants is one effective way to eliminate the risk of invasions from alien plant species. </p>

归化是入侵的前期阶段, 对归化植物的深入研究是科学管理外来物种的基本前提, 有助于预防外来植物造成入侵危害, 区域性归化植物的研究对地方外来植物的管理与防治对策的制定具有重要的指导意义。本文基于文献报道、标本信息和分类学考证, 特别是于2014-2019年对华东地区(安徽省、福建省、江苏省、江西省、山东省、上海市和浙江省)归化植物的全面调查, 统计分析了华东地区归化植物的物种组成和分布格局, 并对其首次引入(或发现)地、引入时间和引入途径进行综合分析。结果表明, 华东地区有归化植物62科181属299种, 总体上呈现种类丰富、以草本植物为主、原产于美洲的种类多、引入途径集中等特征。在空间尺度上, 福建省的物种多样性明显高于其他省市, 且仅分布于该省的种类也最多(达57种, 占总种数的19.1%), 其他省市之间则差别不大。对外交流程度和原产地与归化地之间的气候相似性可能是影响归化植物多样性和空间分布格局的主要因素。在时间尺度上, 1850年之后, 华东地区的归化植物呈现出指数增长的趋势, 增长速率达1.5种/年, 且当前正处于快速增长阶段; 21世纪以来, 80%以上的归化植物来自于无意引入的物种, 这提示我们需要对此特别关注。华东地区作为对外交流的重要区域, 面临着外来植物输入以及归化并造成入侵的双重压力, 因此需对该地区的热点地区如口岸、港口等地加强出入境监管, 同时构建一套行之有效的外来植物风险评估体系, 这是开展外来物种风险管理的基础, 也是预防外来物种入侵的有效手段之一。

We used the literature, field investigations, specimen records and taxonomic research to categorize 515 alien invasive plant species from 72 families and 285 genera reported from China according to their biological and ecological characters, extent of spread within China, and amount of damage caused. Each species was put into one of 5 categories: malignant invasion (34 species), serious invasion (69 species), local invasion (85 species), general invasion (80 species), and species requiring further observation (247 species). According to our analyses of geographic distributional patterns, Chinese alien invasive plants were highly concentrated in the southwestern and eastern coastal provinces of China, and were rapidly spreading to inland China. These categories and distributional patterns of alien invasive plants could help to assess and control their invasive risk. Our results provide baseline information on Chinese alien invasive plants; and, at the same time, we put forward preliminary suggestions for further study, public education and the management of the important alien invasive plants in China.

基于文献报道、野外调查、标本记录和必要的分类学考证, 整理出我国外来入侵植物72科285属515种。根据外来入侵种的生物学与生态学特性、自然地理分布、入侵范围以及所产生的危害, 将其划分为5类: 即恶性入侵类(34种)、严重入侵类(69种)、局部入侵类(85种)、一般入侵类(80种)和有待观察类(247种)。通过地理分布格局分析, 中国外来入侵植物主要分布在我国西南及东部沿海地区, 并进一步扩散到内陆各省。中国外来入侵等级划分和地理分布格局可以作为外来入侵植物风险防范和管理的依据。本工作提供我国外来入侵植物的本底资料的同时, 还对主要外来入侵植物的管理提出了具体建议。

记载了在福建省发现的兰科石斛属植物一新变种——文卉石斛Dendrobium luoi var. wenhuii W. L. Yang，并描述其与原变种罗氏石斛D. luoi L. J. Chen et W. H. Rao和近缘种D. zhenghuoense S. P. Chen, L. Ma & M. H. Li的形态特征区别。

Living gymnosperms comprise four major groups: <i>cycads</i>, <i>Ginkgo</i>, conifers, and gnetophytes. Relationships among/within these lineages have not been fully resolved. Next generation sequencing has made available a large number of sequences, including both plastomes and single-copy nuclear genes, for reconstruction of solid phylogenetic trees. Recent advances in gymnosperm phylogenomic studies have updated our knowledge of gymnosperm systematics. Here, we review major advances of gymnosperm phylogeny over the past 10 years and propose an updated classification of extant gymnosperms. This new classification includes three classes (Cycadopsida, Ginkgoopsida, and Pinopsida), five subclasses (Cycadidae, Ginkgoidae, Cupressidae, Pinidae, and Gnetidae), eight orders (Cycadales, Ginkgoales, Araucariales, Cupressales, Pinales, Ephedrales, Gnetales, and Welwitschiales), 13 families, and 86 genera. We also described six new tribes including Acmopyleae Y. Yang, Austrocedreae Y. Yang, Chamaecyparideae Y. Yang, Microcachrydeae Y. Yang, Papuacedreae Y. Yang, and Prumnopityeae Y. Yang, and made 27 new combinations in the genus <i>Sabina</i>.

— Euphorbia xiangxiui, a new species of Euphorbiaceae from Eastern China, supported by morphological and molecular evidence, is illustrated and described here. It can be distinguished from its closest relatives by the combination of several characters,\n including taller stature (up to 1.6 m), leaves of 5‐18 × 0.6‐3 cm in size, color of primary involucral leaves and cyathophylls changing from pink to rose, fading to green after fruiting, and four to six reniform-rounded concave glands. The molecular phylogenetic analysis,\n based on combined nuclear ribosomal ITS and plastid ndhF sequences, further supports that E. xiangxiui should be placed in the East Asian Clade of Euphorbia subg. Esula sect. Helioscopia.

石松类和蕨类植物是以孢子繁殖的维管植物, 在陆地植物演化上占据重要地位。随着分子系统学研究的开展, 各大类群间的系统发育关系得以阐明, 传统上的概念得以修正, 新的现代石松类和蕨类植物的分类系统也被提出, 并不断得到完善。该文介绍国内外在蕨类植物系统发育方面的研究成果, 重点讨论中国分布的类群的分类处理。文中提出了一个完整的中国现代石松类和蕨类植物的分类系统, 包括5亚纲、14目、39科及12亚科、约140余属。

Since the Fujian Flora published in 1995, more and more new plants have been found in Fujian. In order to master the latest complete data, we collected the related literatures from the database, and analyzed the data from these literatures. The results showed that 4 families and 36 genera 260 species of vascular plants were added on the basis of the Fujian Flora which collecting 248 families and 1596 genera 4724 species of vascular plants (including subspecies and varietas). So the data of vascular plants in Fujian province had increased to 252 families and 1632 genera 4984 species (including subspecies and varietas) by the end of 2014.

自《福建植物志》(1～6卷)出版以来，陆续在福建发现一些植物新种或新记录。文中通过对中外文献数据库进行检索，以发表的论文论著为依据，整理出福建维管束植物新种或新记录种名录和相关数据。截止2014年底，福建省维管束植物在《福建植物志》(1～6卷)收载的248科1596属4724种的基础上，增加了4科36属260种，达252科1632属4984种(含亚、变种)。

Bredia (Melastomataceae) is an Asian genus that extends from central and southern mainland China to Taiwan and the Ryukyu islands. Molecular phylogenetic analyses reveal that the type of Bredia is nested in a clade of 20 species, while Tashiroea, a genus previously synonymized in Bredia, falls in another distantly related clade of 10 species. Our morphological survey shows that the two clades can be distinguished by several diagnostic features including leaf indumentum, texture, leaf surface sculpture under SEM, presence/absence of yellowish uniseriate trichomes, and capsule morphology. Based on molecular and morphological evidence, Bredia is recircumscribed and Tashiroea is resurrected. Description and a list of species are provided for the two genera with the description of a new species, T.villosa.