1 Eastern China Conservation Centre for Wild Endangered Plant Resources, Shanghai Chenshan Botanical Garden, Shanghai 201602, China 

2 Center for Integrative Conservation & Yunnan Key Laboratory for Conservation of Tropical Rainforests and Asian Elephants, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Kunming 650223, China 

3 Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Naypyidaw 05282, Myanmar 

4 State Key Laboratory of Agricultural and Forestry Biosecurity, College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou 350002, China 

5 School of Ecology and Nature Conservation, Beijing Forestry University, Beijing 100083, China