路径依赖下的物种形成机制
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Path-dependent speciation in the process of evolution
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图5. 不同时刻下表型和基因型的频率分布。(a)表示状态量x1(x2)未分化初期某时刻的频率分布; (b)表示群体内的表型x1在分化之后某时刻的频率分布; (c)表示群体内的基因型x2在分化之后相同时刻的频率分布。假设对于原始物种A, 其任意在两个状态x1和x2随着时间的演化受到基因突变和随机漂变的影响, 在相同或不同时刻出现不同程度的分化。当我们处于物种进化的一个“切面”去观察时, 表型x1分化成x11和x12, 并且进化依赖于特定的路径。若此时分化出的表型和基因型在数量统计上满足一定的条件, 同时在表型和基因型两个状态下同时具有特征x11, x21(x11, x22或x12, x21或x12, x22)的个体为新物种。 |
Fig. 5. Frequency distribution of phenotypes and genotypes at different time. (a) showing the frequency distribution curve of the state x1(x2) in the early differentiation. (b) demonstrating the frequency distribution curve of the phenotype x1 at a certain moment after differentiation. (c) indicating the frequency distribution curve of the genotype x2 at the same moment after differentiation. Suppose that any two states x1 and x2 of the original species A evolved influenced by random mutation and drift, then different degree of differentiation occurs in the same or different time. When observing in a “cutting plane” of a species evolution, we notice that x1(x2) differentiates into x11(x21) and x12(x22), and the evolution are path-dependent. Meanwhile, if the differentiated phenotypes and the genotypes in a quantity statistics satisfy certain conditions, the individuals with ${{x}_{11}},{{x}_{21}}({{x}_{11}},{{x}_{22}},or{{x}_{12}},{{x}_{21}},or{{x}_{12}},{{x}_{22}})$ are new species, respectively. |
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