生物多样性 ›› 2003, Vol. 11 ›› Issue (4): 295-302.  DOI: 10.17520/biods.2003037

• 论文 • 上一篇    下一篇

长白山阔叶红松林结构多样性的初步研究

郑景明,罗菊春   

  1. 1 (中国科学院植物研究所植被数量生态学重点实验室,北京 100093)
    2 (北京林业大学,北京 100083)
  • 收稿日期:2002-12-12 修回日期:2003-06-09 出版日期:2003-07-20 发布日期:2003-07-20
  • 通讯作者: 郑景明

Structural diversity of broadleaved Korean pine forest in Changbai Mountain

ZHENG Jing-Ming, LUO Ju-Chun   

  1. 1 laboratory of Quantitative Vegetation Ecology,Institute of Botany,Chinese Academy of Siciences,Beijing 100093
    2 Beijing Forestry University,Beijing 100083
  • Received:2002-12-12 Revised:2003-06-09 Online:2003-07-20 Published:2003-07-20
  • Contact: ZHENG Jing-Ming

摘要: 在林业可持续发展的背景下,对森林实施生态系统管理,必须保护和维持森林的生物多样性。这就要求在森林调查和经营过程中增加林分结构多样性方面的内容,因此必须充分认识森林结构包括的内容,并研究制订简便实用的结构多样性指标。本文采用目测分层盖度结合无样地点四分法取样进行林分结构因子调查,设计了一套简便的林分结构指标体系,并对长白山红松阔叶混交林及其次生林进行了林分结构多样性的测定研究。结果表明:基于分层盖度构建的林分垂直多样性指数和水平结构异质性指数,可以较好地表示林分的基本结构特征,同时借助点四分法测定的物种多样性指数和对倒木、站干、林隙等的统计,可以较全面地描述阔叶红松林的林分结构因素。应用该方法测定不同的原始林样地的林分结构,得到的林分结构指标与其他林分结构因素的描述,同常规林学调查结论一致并更全面;同时对不同采伐方式的林分结构进行了测定,结合相应的常规指标测定对比,表明该结构指数不但能较好地表示群落的空间结构特征,同时也能对比不同经营方式对林分结构造成的影响,如择伐方式比皆伐方式对林分的结构影响相对较小而且结构恢复时间较短,从而说明对阔叶红松林这种复层异龄混交林的森林类型而言,择伐是一种更适合的通过结构调控进行生态系统管理的方式。在对生态系统进行经营管理过程中,采用该方法能有效地说明林分的垂直结构复杂性和水平结构异质性及重要的林分结构因子存在状况,因而更适合当前森林生态系统管理的要求。文章进一步分析了林分结构指标待改进的部分与研究方向。

AbstractFrom the background of sustainable development of forestry, biodiversity in forest should be maintained and protected when managing an ecosystem. However, as a result of the complex meaning of biodiversity, ordinary indices used in ecology and forestry are not commonly used by forest managers because of their high requirements of data and precision in order to convey so much information on biodiversity in a single index. On the basis of recent research on virgin forests around the world, Franklin advocates that diversity of forest stand structure could be used as an indirect indicator of forest biodiversity, and this has been accepted by many forest experts. As a result, some new ideas on how to manage forest ecosystems through structure control or restructuring have been put forward. In order to manage forest ecosystem through stand structure control, the significance of forest structure must be clarified and some new indices must be created for use in forest survey and in proper forest measurements. In this study, a set of structural diversity indices for broadleaved Korean pine forest were put forward and tested in different types of stands, including virgin forest, forest after selection cutting, and forest regenerated after clear cutting. Based on the measurement of stratified coverage in addition to plotless sampling, the vertical structure index (VSI ) and horizontal heterogeneity index ( HHI ) were generated, along with descriptions of other structural elements such as downed logs, standing poles and canopy gaps. The basic meaning of VSI is the volume of space occupied by branches and leaves, which was calculated by coverage in each layer and its weight. The basic meaning of HHI is the difference of coverage in all layers between locations in the stands, which was calculated as a community dissimilarity index. The bigger the two indices, the more habitat types for living organisms exist in the forest. At the same time, other ancillary indices, such as species composition, amount of coarse woody debris and gaps were surveyed through plotless sampling methods and transects with plots. Comparison between different stands of virgin forest indicates that these indices were precise enough to describe the spatial structure of the stands and were in agreement with analyses resulting from common forestry survey methods. Used in secondary forest stands, they could also provide more information than common single biodiversity indices. Before using data from plotless sampling, the similarity coefficients between plot sampling and plotless sampling were calculated. The precision of the two methods proved to be identical. Used in stands with different disturbance histories, these indices illustrated remarkable differences in the structures of the stands, indicating that the indices are suitable for forest survey and for guiding choice of management measures in forest ecosystem management. Disadvantages and methods for improvement of the indices are discussed and research priorities are proposed.